Heterotheca grandiflora |
Heterotheca villosa |
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silk-grass goldenaster, telegraph weed |
golden-aster, hairy false goldenaster, hairy goldaster, hairy golden-aster |
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Habit | Annuals, biennials, or short-lived perennials, 10–250 cm (atypical, spring-blooming, dwarf dune plants have short stems; perennating from proximal stem nodes in south; aromatic); taprooted. | Perennials, (5–)16–40(–70) cm; taprooted. | ||||||||||||||||||||||||||||||||
Stems | 1–5+, ascending to erect (strict, 0.9–1.7 cm diam., only distal lateral branches developing in larger plants), moderately to densely hispido-strigose, more so proximally, moderately to densely stipitate-glandular, more so distally. |
1–50+, decumbent to erect (sometimes brown or reddish brown, sometimes whitish distally, sometimes ± brittle), sparsely to densely hispido-strigose, sparsely to abundantly long-hispid, eglandular or sparsely to densely stipitate-glandular. |
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Leaves | basal and proximal cauline petiolate (petioles 3–7 cm, bases auriculate-clasping), blades ovate or elliptic to oblong, 50–80 × 20–30 mm, margins flat or somewhat undulate, coarsely serrate to serrulate or entire, proximally long-ciliate, apices acute, faces moderately to densely long-hispido-strigose; distal sessile, blades oblong-lanceolate to lanceolate, 20–40 × 20–30 mm, reduced distally, margins often undulate, coarsely serrate. |
generally not crowded; proximal cauline petiolate, blades oblanceolate, (90–)220–400(–600) × (2–)4–8(–13) mm, bases cuneate, margins flat, usually entire, rarely with 1–2 apical teeth, strigoso-ciliate, sparsely to abundantly long-hispido-strigose proximally, apices acute to obtuse, sometimes mucronate, faces sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular; distal sessile, blades usually lanceolate or oblanceolate to oblong, rarely ovate or lanceolate-triangular, (4–)15–28(–42) × (1.5–)3.5–7(–12.5) mm, bases attenuate to convex-cuneate to rounded, margins usually flat, rarely remotely undulate, strigoso-ciliate, sparsely to abundantly long-hispido-strigose proximally, apices acute to obtuse, sometimes mucronate, faces sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular. |
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Peduncles | 10–65 mm, sparsely to densely hispido-strigose, densely stipitate-glandular; bracts 0–3, linear-lanceolate to linear, reduced distally. |
(4–)15–41(–98) mm, sparsely to densely hispido-canescent, eglandular or sparsely to densely stipitate-glandular; bracts 1–7+, usually linear-oblanceolate, rarely leaflike and linear-oblanceolate, usually greatly reduced, margins usually flat, rarely remotely undulate, strigoso-ciliate, sparsely to abundantly long-hispido-strigose proximally, apices acute, sometimes mucronate, faces sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular; rarely 1–2 leaflike, oblanceolate bracts subtending heads. |
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Involucres | campanulo-hemispheric, 6–9 mm. |
narrowly cylindric to campanulate, (5–)6–9.5(–13) mm. |
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Ray florets | 20–40; laminae 5–8 × 1–2 mm. |
(5–)10–20(–38); laminae (3.5–)6.5–11(–20) × 1–2(–3) mm. |
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Disc florets | 30–75; corollas 4–6 mm, lobes 0.5–0.8 mm. |
(10–)20–50(–85); corollas barely ampliate, (4–)5–6(–8) mm, glabrous or glabrate (few, minute hairs), lobes 0.4–0.75(–1) mm, glabrous or glabrate (hairs 0.1–0.35 mm). |
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Phyllaries | in 4–6 series, linear-lanceolate, unequal, margins scarious, faces strigose, densely stipitate-glandular. |
in 4–5 series, lanceolate to linear-lanceolate or triangular-lancelate, unequal (outer 1/5–1/3 length of inner), margins scarious, sometimes reddish purple distally, ciliate distally or apically, faces sparsely to densely strigose, eglandular or sparsely to moderately stipitate-glandular. |
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Heads | 10–110+, in corymbiform arrays, becoming paniculiform with age, branches ascending, usually relatively short compared to plant height. |
1–16(–42), in usually open, corymbiform, rarely paniculiform arrays. |
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Cypselae | dimorphic, (ray) 2–5 mm, faces glabrous or slightly strigose on angles, (disc) 4–6 mm, faces moderately to densely strigose; pappi 0 (ray) or (disc) stramineous to rusty brown, outer bristles 0.4–0.7 mm, inner 6–9 mm, longest weakly clavate. |
monomorphic, obconic, compressed, 1.7–2.7(–3.4) mm, ribs 4–8(–10) (rarely brownish), faces sparsely to moderately strigose; pappi off-white, outer of linear scales 0.25–1 mm, inner of 30–45 bristle (4–)5–6.5(–8.5) mm, longest weakly clavate (usually equaling or longer than corollas). |
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2n | = 18. |
= 18, 36. |
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Heterotheca grandiflora |
Heterotheca villosa |
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Phenology | Flowering Apr–Dec(–Jan). | |||||||||||||||||||||||||||||||||
Habitat | Sandy and gravelly soils, dunes, disturbed ground, roadsides, vacant lots, fields | |||||||||||||||||||||||||||||||||
Elevation | 0–100(–900) m (0–300(–3000) ft) | |||||||||||||||||||||||||||||||||
Distribution |
AZ; CA; NV; UT [Introduced in Pacific Islands (Hawaii)]
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AZ; CA; CO; IA; ID; IL; KS; MI; MN; MT; ND; NE; NM; NV; OR; SD; TX; UT; WA; WI; WY; AB; BC; MB; ON; SK
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Discussion | Heterotheca grandiflora is common in California, rare and probably introduced in Arizona, Nevada, and Utah, and introduced in Hawaii. Rosette plants can bloom in the spring. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 9 (9 in the flora). Heterotheca villosa is variable in stem height, leaf base shape, stem and leaf indument, number of heads, and number of florets. Thus, the species is difficult to circumscribe because each variety has a suite of diagnostic traits and a distribution that is restricted to a portion of the range of the species. Generally, var. minor (glandular), and to a lesser extent, var. foliosa (eglandular) are the glue holding the other varieties together in a widely distributed polymorphic species; this is comparable to the situation in H. sessiliflora, in which var. echioides is the glue. Variety minor hybridizes with all other varieties. Numerous local races occur that are sometimes quite distinct when extreme, but they intergrade with one or more other races, especially in var. minor (J. C. Semple 1996). The species is divided here on the basis of indument features, leaf shape, and stem height, paralleling the infraspecific treatments of H. fulcrata and H. sessiliflora. H. A. Gleason and A. Cronquist (1991) and Cronquist (1994) acknowledged the variability of the species and the existence of many local races, but lumped most of these into var. villosa and var. hispida of Chrysopsis villosa, in which they also included H. camporum, H. canescens, H. fulcrata, H. pumila, H. stenophylla var. angustifolia, H. viscida, and H. zionensis. Diploid races are usually distinct from each other, but each has given rise to one or more tetraploid lines that are less distinct. The treatment here is based on the detailed presentation in Semple. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 20, p. 235. | FNA vol. 20, p. 2. | ||||||||||||||||||||||||||||||||
Parent taxa | Asteraceae > tribe Astereae > Heterotheca | Asteraceae > tribe Astereae > Heterotheca | ||||||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||||||||||
Synonyms | H. floribunda | Amellus villosus, Chrysopsis villosa, Diplogon villosum, Diplopappus villosus | ||||||||||||||||||||||||||||||||
Name authority | Nuttall: Trans. Amer. Philos. Soc., n. s. 7: 315. (1840) | (Pursh) Shinners: Field & Lab. 19: 71. (1951) | ||||||||||||||||||||||||||||||||
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