Heterotheca grandiflora |
Heterotheca subaxillaris |
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silk-grass goldenaster, telegraph weed |
camphor false golden-aster, camphorweed, golden aster, unknown |
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Habit | Annuals, biennials, or short-lived perennials, 10–250 cm (atypical, spring-blooming, dwarf dune plants have short stems; perennating from proximal stem nodes in south; aromatic); taprooted. | Annuals or biennials, 10–200 cm (aromatic); taprooted (rarely perennating in south from proximal stem nodes). | ||||
Stems | 1–5+, ascending to erect (strict, 0.9–1.7 cm diam., only distal lateral branches developing in larger plants), moderately to densely hispido-strigose, more so proximally, moderately to densely stipitate-glandular, more so distally. |
1–4+, procumbent to erect (sometimes reddish brown, proximal to distal branches well developed in larger plants), sparsely to densely hispido-strigose, sparsely to densely stipitate-glandular. |
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Leaves | basal and proximal cauline petiolate (petioles 3–7 cm, bases auriculate-clasping), blades ovate or elliptic to oblong, 50–80 × 20–30 mm, margins flat or somewhat undulate, coarsely serrate to serrulate or entire, proximally long-ciliate, apices acute, faces moderately to densely long-hispido-strigose; distal sessile, blades oblong-lanceolate to lanceolate, 20–40 × 20–30 mm, reduced distally, margins often undulate, coarsely serrate. |
basal and proximal rarely persisting to flowering, sometimes present and withered, brown to black; basal to mid cauline petiolate (petioles 10–40 mm, bases auriculate-clasping), blades ovate to elliptic or lanceolate, 10–70 × 6–55 mm, bases cuneate to attenuate, margins flat or undulate, coarsely serrate or entire, basally long-ciliate, apices acute, faces moderately hispido-scabrous; distal sessile, blades ovate to lanceolate, 10–90 × 2–20 mm, reduced distally, bases often becoming cordate distally, subclasping or not clasping, margins entire, faces sparsely to moderately hispido-strigose, sparsely to densely stipitate-glandular. |
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Peduncles | 10–65 mm, sparsely to densely hispido-strigose, densely stipitate-glandular; bracts 0–3, linear-lanceolate to linear, reduced distally. |
2–35 mm, sparsely to densely hispido-strigose, moderately to densely stipitate-glandular; bracts 0–4, proximal leaflike, ovate to lanceolate, reduced distally and becoming linear, faces sparsely to moderately hispido-strigose, sparsely to densely stipitate-glandular. |
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Involucres | campanulo-hemispheric, 6–9 mm. |
hemispheric to campanulate, 4–8(–10) mm. |
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Ray florets | 20–40; laminae 5–8 × 1–2 mm. |
15–35; laminae 3–7(–9) × 1–2 mm. |
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Disc florets | 30–75; corollas 4–6 mm, lobes 0.5–0.8 mm. |
25–60; corollas weakly ampliate, 2–9 mm, proximal throats glabrous to sparsely short-strigose, lobes 0.5–0.7 mm, glabrous. |
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Phyllaries | in 4–6 series, linear-lanceolate, unequal, margins scarious, faces strigose, densely stipitate-glandular. |
in 4–6 series, lanceolate, sometimes strongly unequal, margins scarious and distally strigoso-ciliate, faces sparsely to densely stipitate-glandular, with 0–28 coarse, scabro-strigose hairs distally. |
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Heads | 10–110+, in corymbiform arrays, becoming paniculiform with age, branches ascending, usually relatively short compared to plant height. |
3–180+ in corymbo-paniculiform arrays, sometimes becoming profusely branched (loosely paniculiform or broadly corymbiform in larger plants), branches ascending to spreading. |
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Cypselae | dimorphic, (ray) 2–5 mm, faces glabrous or slightly strigose on angles, (disc) 4–6 mm, faces moderately to densely strigose; pappi 0 (ray) or (disc) stramineous to rusty brown, outer bristles 0.4–0.7 mm, inner 6–9 mm, longest weakly clavate. |
dimorphic, obconic, ribs 2–3, (ray) triangular in cross section, 1.5–2.5 mm, faces glabrous to slightly strigose, (disc) laterally compressed, 2–4 mm, faces moderately to densely strigose; pappi 0 (ray) or (disc) tan to rust, outer of linear to triangular scales 0.25–0.6 mm, inner of 25–45 bristles 4–9 mm, longest weakly clavate. |
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2n | = 18. |
= 18. |
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Heterotheca grandiflora |
Heterotheca subaxillaris |
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Phenology | Flowering Apr–Dec(–Jan). | |||||
Habitat | Sandy and gravelly soils, dunes, disturbed ground, roadsides, vacant lots, fields | |||||
Elevation | 0–100(–900) m (0–300(–3000) ft) | |||||
Distribution |
AZ; CA; NV; UT [Introduced in Pacific Islands (Hawaii)]
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AL; AR; AZ; CA; CO; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MD; MO; MS; NC; NE; NJ; NM; NV; NY; OK; PA; SC; TN; TX; UT; VA; Mexico
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Discussion | Heterotheca grandiflora is common in California, rare and probably introduced in Arizona, Nevada, and Utah, and introduced in Hawaii. Rosette plants can bloom in the spring. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). Heterotheca subaxillaris is a very weedy, morphologically variable species that has been treated as three species (B. L. Wagenknecht 1960; V. L. Harms 1965, 1970; J. C. Semple 1996) or as a single polymorphic species (G. L. Nesom 1990e). Harms (1965) acknowledged the difficulty in finding consistent characters to separate plants on a geographic basis, as did Nesom (1990e). Harms (1970) noted that H. psammophila and H. latifolia are probably conspecific. It seems best to treat the complex as a single polymorphic species. In historic times, human-assisted dispersal of possibly once distinct races has likely resulted in blurring of such distinctions with one exception. Plants of the outer coastal plain from Texas to North Carolina usually have a cluster of coarse hairs near the tip of phyllaries, at least in summer and fall-blooming plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 20, p. 235. | FNA vol. 20, p. 234. | ||||
Parent taxa | Asteraceae > tribe Astereae > Heterotheca | Asteraceae > tribe Astereae > Heterotheca | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | H. floribunda | Inula subaxillaris | ||||
Name authority | Nuttall: Trans. Amer. Philos. Soc., n. s. 7: 315. (1840) | (Lamarck) Britton & Rusby: Trans. New York Acad. Sci. 7: 10. (1887) | ||||
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