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silk-grass goldenaster, telegraph weed

stiffleaf false goldenaster, stiffleaf goldenaster

Habit Annuals, biennials, or short-lived perennials, 10–250 cm (atypical, spring-blooming, dwarf dune plants have short stems; perennating from proximal stem nodes in south; aromatic); taprooted. Perennials 18–46(–65) cm; taprooted, sometimes spreading by rhizomes, forming additional clumps.
Stems

1–5+, ascending to erect (strict, 0.9–1.7 cm diam., only distal lateral branches developing in larger plants), moderately to densely hispido-strigose, more so proximally, moderately to densely stipitate-glandular, more so distally.

1–50+, ascending to erect (sometimes brown, or reddish to dark brown, often proximally brittle), often short-branched in distal 1/2, sparsely to densely long-hispid and strigose (hairs often broken off, especially proximally), sometimes becoming moderately hairy and densely stipitate-glandular distally (axillary fascicles of leaves often present).

Leaves

basal and proximal cauline petiolate (petioles 3–7 cm, bases auriculate-clasping), blades ovate or elliptic to oblong, 50–80 × 20–30 mm, margins flat or somewhat undulate, coarsely serrate to serrulate or entire, proximally long-ciliate, apices acute, faces moderately to densely long-hispido-strigose;

distal sessile, blades oblong-lanceolate to lanceolate, 20–40 × 20–30 mm, reduced distally, margins often undulate, coarsely serrate.

generally ascending, congested;

proximal cauline petiolate, blades linear-oblanceolate, (15–)24–42 × (2–)3–6(–8.5) mm, bases cuneate to attenuate, margins flat, entire, strigoso-ciliate, proximally long-hispido-strigose, apices acute, faces sparsely to moderately hispido-strigose and sparsely to moderately stipitate-glandular, or moderately to densely hispido-strigose and eglandular;

distal sessile (bright green to grayish green), blades linear-oblanceolate to narrowly oblanceolate (rarely broader), (9–)16.5–30 × (1.5–)2.2–4.3(–6) mm, usually little reduced distally (often surpassing heads), bases cuneate to attenuate, margins flat, often abundantly long-hispido-strigose along whole, apices acute, sometime mucronulate to white-spinulose, faces sparsely to moderately densely hispido-strigose (2–65 hairs/mm²; hairs often scabrous, ± pustulate), usually eglandular, sometimes sparsely to densely stipitate-glandular.

Peduncles

10–65 mm, sparsely to densely hispido-strigose, densely stipitate-glandular;

bracts 0–3, linear-lanceolate to linear, reduced distally.

2–16 mm, sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular;

bracts 1–3, proximal linear-oblanceolate to oblanceolate, sometimes little reduced distally, leaflike.

Involucres

campanulo-hemispheric, 6–9 mm.

cylindro-turbinate to narrowly campanulate, (4.5–)5.5–8(–11) mm.

Ray florets

20–40;

laminae 5–8 × 1–2 mm.

10–24(–36);

laminae (5–)7–11.5(–16.5) × 0.7–1.5(–2.1) mm.

Disc florets

30–75;

corollas 4–6 mm, lobes 0.5–0.8 mm.

(12–) 23–55(–70);

corollas slightly ampliate, 4.5–7 mm, throats glabrous, lobes 0.4–0.9 mm, lobes sparsely pilose (hairs on lobes 0.1–0.25 mm).

Phyllaries

in 4–6 series, linear-lanceolate, unequal, margins scarious, faces strigose, densely stipitate-glandular.

in 5–6 series, unequal (outer 1/5–1/4 length of inner), narrowly triangular-lanceolate to lanceolate, margins scarious, faces sparsely to moderately strigose, sparsely to densely stipitate-glandular.

Heads

10–110+, in corymbiform arrays, becoming paniculiform with age, branches ascending, usually relatively short compared to plant height.

(1–)2–16(–30), in corymbiform arrays, branches ascending.

Cypselae

dimorphic, (ray) 2–5 mm, faces glabrous or slightly strigose on angles, (disc) 4–6 mm, faces moderately to densely strigose;

pappi 0 (ray) or (disc) stramineous to rusty brown, outer bristles 0.4–0.7 mm, inner 6–9 mm, longest weakly clavate.

monomorphic, obconic, compressed, 1.5–2.5(–3) mm, ribs 6–10 (sometimes brownish), faces sparsely to moderately strigose;

pappi off-white, outer of linear scales 0.25–0.5 mm, inner of 25–40 bristles (4–)5–6.5(–7.5) mm, longest weakly clavate.

2n

= 18.

= 18, 36.

Heterotheca grandiflora

Heterotheca stenophylla

Phenology Flowering Apr–Dec(–Jan).
Habitat Sandy and gravelly soils, dunes, disturbed ground, roadsides, vacant lots, fields
Elevation 0–100(–900) m (0–300(–3000) ft)
Distribution
from FNA
AZ; CA; NV; UT [Introduced in Pacific Islands (Hawaii)]
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; IA; KS; NE; NM; OK; SD; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Heterotheca grandiflora is common in California, rare and probably introduced in Arizona, Nevada, and Utah, and introduced in Hawaii. Rosette plants can bloom in the spring.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Heterotheca stenophylla is divided into two varieties that differ in gland and hair density. Their ranges overlap to a great extent, and intermediates occur throughout the range. Both H. stenophylla and H. canescens often have numerous, overlapping leaves and often bear axillary fascicles of leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Distal mid stems glabrate to sparsely hispido-strigose, moderately to densely stipitate- glandular; leaf faces sparsely to moderately hispido-strigose, moderately to densely stipitate-glandular (appearing dark to bright green), hair bases often broad (pustulate); phyllary faces usually sparsely strigose and stipitate-glandular
var. stenophylla
1. Distal mid stems moderately to densely hispido-strigose, eglandular or sparsely glandular; leaf faces moderately to densely hispido-strigose, eglandular or sparsely stipitate-glandular beneath hairs (appearing pale green to gray-green), hairs rarely broad-based, pustulate; phyllary faces usually moderately strigose, eglandular
var. angustifolia
Source FNA vol. 20, p. 235. FNA vol. 20, p. 246.
Parent taxa Asteraceae > tribe Astereae > Heterotheca Asteraceae > tribe Astereae > Heterotheca
Sibling taxa
H. camporum, H. canescens, H. fulcrata, H. jonesii, H. marginata, H. monarchensis, H. oregona, H. pumila, H. rutteri, H. sessiliflora, H. shevockii, H. stenophylla, H. subaxillaris, H. villosa, H. viscida, H. zionensis
H. camporum, H. canescens, H. fulcrata, H. grandiflora, H. jonesii, H. marginata, H. monarchensis, H. oregona, H. pumila, H. rutteri, H. sessiliflora, H. shevockii, H. subaxillaris, H. villosa, H. viscida, H. zionensis
Subordinate taxa
H. stenophylla var. angustifolia, H. stenophylla var. stenophylla
Synonyms H. floribunda Chrysopsis hispida var. stenophylla, Chrysopsis stenophylla, Chrysopsis villosa var. stenophylla
Name authority Nuttall: Trans. Amer. Philos. Soc., n. s. 7: 315. (1840) (A. Gray) Shinners: Field & Lab. 19: 68. (1951)
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