Heteromeles |
Rosaceae subfam. amygdaloideae |
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Christmas berry, toyon |
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Habit | Shrubs or trees, canopies dense, 20–100 dm. | Shrubs or trees, sometimes subshrubs or herbs. |
Stems | usually 1; bark (trunk) grayish, with fine tan to dark gray striations, ± smooth; short shoots absent; unarmed; young stems puberulent. |
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Leaves | persistent, cauline, simple; stipules variably persistent, free, minute, margins unknown, apex gland-tipped; petiole present; blade ± elliptic to narrowly elliptic or narrowly oblong, 5–15(–20) cm, leathery, margins revolute, sharply, remotely serrate, venation pinnate (craspedodromous-brochidodromous), surfaces tomentose when young, glabrescent. |
alternate, sometimes opposite, simple, sometimes pinnately compound; stipules present or absent. |
Inflorescences | terminal, 20–150-flowered, panicles, ± dome-shaped, white-tomentose; bracts present on proximal nodes, leaflike, plus numerous, scalelike appendages on axes; bracteoles present caducous, delicate. |
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Pedicels | present. |
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Flowers | perianth and androecium epigynous, 10 mm diam.; hypanthium urceolate, 2–4 mm, glabrous or weakly floccose; sepals 5, suberect, triangular; petals 5, white, irregularly round, base weakly clawed; stamens 10, shorter than petals; carpels 2 or 3, distinct, basally adnate to hypanthium, styles 2 or 3, lateral, distinct; ovules 2. |
torus absent or minute; carpels 1–5(–8), distinct or +/- connate (Maleae), free or +/- adnate to hypanthium (many Maleae), styles distinct or +/- connate (some Maleae); ovules (1 or)2(–5+), collateral, clustered, or biseriate. |
Fruits | pomes, usually bright red, sometimes yellow, ellipsoid, 5–10 mm, glabrous or glabrate; flesh mealy; hypanthium persistent; sepals persistent, accrescent over hypanthial opening. |
follicles aggregated or not, capsules, drupes aggregated or not, aggregated drupelets, pomes, or aggregated nutlets, rarely achenes or aggregated achenes; styles persistent or deciduous, not elongate (elongate in Gillenieae). |
Pyrenes | 2 or 3 per fruit, carpel walls thin, soft; styles not persistent; seeds 1 per pyrene, large. |
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x | = 8, 9, 15, 17. |
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Heteromeles |
Rosaceae subfam. amygdaloideae |
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Distribution |
CA; nw Mexico |
HI; North America; Mexico; Central America; South America; Europe; Asia; Africa; Atlantic Islands (Madeira); Australia |
Discussion | Species 1. Heteromeles resembles Aronia and Photinia; it differs by having 10 stamens instead of 20, nearly free carpels (versus fully connate), and fruit flesh lacking stone cells (versus often possessing some). Its fruit is of the coreless type (J. R. Rohrer et al. 1991). Molecular evidence (D. Potter et al. 2007) shows a relationship to the Asiatic Eriobotrya and Rhaphiolepis Lindley miniclade; fruit type, notched petals, and other characteristics agree. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cyanogenic glycosides are usually present in Amygdaloideae; sorbitol is present. The name Amygdaloideae Arnott (1832) has priority over Spiraeoideae Arnott (1832), used by D. Potter et al. (2007), because Amygdalaceae (1820) is an earlier conserved name. Tribes 9, genera 55, species ca. 1300 (9 tribes, 38 genera, 361 species, including 20 hybrids, in the flora) (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 447. | FNA vol. 9, p. 345. |
Parent taxa | ||
Subordinate taxa | ||
Name authority | M. Roemer: Fam. Nat. Syn. Monogr. 3: 100, 105. (1847) | Arnott: Botany, 107. (1832) |
Web links |