Flora of North America species comparison

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Hesperidanthus

Hesperidanthus barnebyi
Habit Perennials or subshrubs; (caudex well-developed, woody); not scapose; (glaucous), glabrous. Perennials.
Stems

(simple or few to several from base), erect or ascending, usually branched distally.

few to several from caudex, erect, 1–3.5(–3.8) dm.
Leaves

cauline;

not rosulate, petiolate, subsessile, or sessile, blade (somewhat fleshy, becoming leathery when dry, base not auriculate), margins entire, denticulate, or coarsely dentate.

subsessile or petiolate, (0.4–1 cm);

blade oblong or elliptic to oblanceolate, 1.5–5 cm × 4–24 mm, base cuneate, margins entire or obscurely denticulate, apex obtuse to rounded.
Racemes

(corymbose, few- to several-flowered, sometimes bracteate proximally), elongated in fruit.

usually 2–8-flowered (rarely more).
Flowers

sepals ascending to erect, oblong, lateral pair usually not saccate basally (slightly saccate in H. linearifolius, apex of median pair cucullate or not);

petals white, lilac, lavender, purple, or yellow (sometimes with darker veins), spatulate, claw differentiated or not from blade;

stamens tetradynamous or subequal;

filaments not dilated basally;

anthers oblong to linear, (apex obtuse or apiculate);

nectar glands: lateral annular or lunar, median glands often confluent with lateral.

sepals green to purple, 5–8 × 2–3 mm;

petals white or lilac (with darker purple veins), 9.5–12 × 2.5–3.5 mm, claw distinctly differentiated from blade, (narrower);

filaments 2.5–3 mm;

anthers linear, 2.5–3 mm;

gynophore 0–1.5 mm.
Fruiting pedicels

divaricate to ascending or suberect, slender.

ascending to divaricate, straight, 10–27 mm.
Fruits

sessile or stipitate, linear, not torulose, terete or slightly latiseptate;

valves each with prominent midvein;

replum rounded;

septum complete;

ovules 8–110 per ovary;

style distinct;

stigma capitate or conical, entire or 2-lobed (lobes prominent, connivent).

straight or curved, terete, 3.5–5.5 cm × 1–2 mm;

ovules 24–42 per ovary;

style subclavate, 1–2 mm;

stigma flat, obscurely 2-lobed.
Seeds

uniseriate, plump, not winged, oblong;

seed coat not or slightly mucilaginous when wetted;

cotyledons incumbent or obliquely so.

1.8–2.2 × 0.9–1.2 mm.
x

= 11.

Hesperidanthus

Hesperidanthus barnebyi
Phenology Flowering Apr–Jun.
Habitat Mixed desert communities
Elevation 1700-2000 m (5600-6600 ft)
Distribution
w United States; n Mexico
[BONAP county map]
from FNA
UT
[BONAP county map]
Discussion

Species 5 (5 in the flora).

For a detailed account on the generic limits and affinities of Hesperidanthus, see I. A. Al-Shehbaz (2005). As delimited here, the genus includes what R. C. Rollins (1993) assigned to Schoenocrambe Greene (minus the type), Caulostramina, and Glaucocarpum. One species is endemic to Inyo County, California, three are highly localized endemics in Utah, and the fifth is widespread in the southwestern United States and northern Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

The species is restricted to the Chinle Formation in Emery and Wayne counties.

Hesperidanthus barnebyi, as Schoenocrambe barnebyi, is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Key
1. Petals yellow; fruits 1-2 cm; ovules 8-16 per ovary.
H. suffrutescens
1. Petals purple, lilac, lavender, or white; fruits (1.8-)2.5-11 cm; ovules 26-110 per ovary
→ 2
2. Median sepals cucullate; stigmas conical, lobes connivent; petal veins not darker than blade; ovules (76-)80-110 per ovary; Arizona, Colorado, New Mexico, Texas.
H. linearifolius
2. Median sepals not cucullate; stigmas flat, lobes entire or obscure; petal veins darker than blade; ovules 26-62 per ovary; California, Utah
→ 3
3. Leaves petiolate, blades ovate to broadly so, 10-35 mm wide, margins coarsely and irregularly dentate; California.
H. jaegeri
3. Leaves sessile, subsessile, or petiolate, blades linear, linear-lanceolate, oblong, elliptic, or oblanceolate, (0.8-)1-24 mm wide, margins entire or obscurely denticulate; Utah
→ 4
4. Leaves sessile or subsessile, blades linear or linear-lanceolate; sepals 4.2-6.5 mm; anthers oblong, 1-1.5 mm.
H. argillaceus
4. Leaves petiolate, blades oblong, elliptic, or oblanceolate; sepals 5-8 mm; anthers linear, 2.5-3 mm.
H. barnebyi
Source FNA vol. 7, p. 689. Author: Ihsan A. Al-Shehbaz. FNA vol. 7, p. 691.
Parent taxa Brassicaceae > tribe Thelypodieae Brassicaceae > tribe Thelypodieae > Hesperidanthus
Sibling taxa
H. argillaceus, H. jaegeri, H. linearifolius, H. suffrutescens
Subordinate taxa
H. argillaceus, H. barnebyi, H. jaegeri, H. linearifolius, H. suffrutescens
Synonyms Thelypodium subg. H., Caulostramina, Glaucocarpum Thelypodiopsis barnebyi, Schoenocrambe barnebyi
Name authority (B. L. Robinson) Rydberg: Bull. Torrey Bot. Club 34: 433. (1907) (S. L. Welsh & N. D. Atwood) Al-Shehbaz: Harvard Pap. Bot. 10: 50. (2005)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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