Herzogiella striatella |
Hypnaceae |
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herzogiella moss, tassel moss |
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Habit | Plants in thin to dense mats, yellowish, dark green, or brownish. | Plants small to large, lax to dense, in tufts or mats, dark green, yellow-green, golden green, or orange, sometimes reddish brown or nearly black, often lustrous. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stem(s) | 2 cm, 0.5–2 mm wide, ascending to erect. |
and branch leaves similar or less commonly differentiated, straight to homomallous, often falcate-secund, usually ovate-lanceolate, often asymmetric, sometimes linear, lanceolate, or triangular, occasionally plicate; base sometimes decurrent; margins often plane, occasionally recurved proximally or throughout, entire or toothed; apex obtuse to acuminate; costa double, short, to obscure or ecostate; alar cells usually differentiated, often quadrate to short-rectangular, sometimes enlarged and inflated, pigmented or similar in color to other cells, excavate or plane; medial and distal laminal cells linear, hexagonal, or elongate-sinuate, smooth, sometimes prorulose at distal ends on abaxial surface. |
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Leaves | loosely imbricate to somewhat spreading, often squarrose, sometimes squarrose-recurved, usually straight at stem and branch apices, not plicate, 0.6–2 × 0.3–0.8 mm; base distinctly decurrent; margins serrulate to strongly serrate; alar cells rounded to oval, abruptly inflated, 14–65 × 14–24 µm, hyaline, sometimes orange to red; basal laminal cell walls pitted, sometimes pitted to mid leaf or beyond; medial cells 24–50 × 4–7 µm. Sexual condition autoicous. |
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Seta | light brown to red, 0.9–2 cm. |
elongate, smooth (occasionally scabrous near capsule in Ctenidium). |
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Sexual condition | autoicous, dioicous, or phyllodioicous. |
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Capsule | inclined to nearly erect, light brown, oblong to cylindric, slightly arcuate to straight, 1–2 × 0.3–0.5 mm, not or little contracted below mouth when dry; operculum conic to conic-apiculate, 0.3–0.4 mm. |
inclined, horizontal, or sometimes erect, cylindric or ovoid, usually smooth, often constricted below mouth when dry and empty; operculum conic or rostrate; peristome usually double; exostome teeth 16, external surface cross striate basally, papillose distally, sometimes bordered, internal surface often trabeculate; endostome usually free, sometimes fused to exostome, basal membrane high or rarely low, segments 16, cilia 1–3, nodose, rarely rudimentary or absent. |
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Calyptra | cucullate, smooth (weakly prorulose distally in Ctenidium), naked or rarely hairy. |
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Spores | 10–15 µm. |
spheric to ovoid, usually finely papillose, rarely smooth. |
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Specialized | asexual reproduction sometimes by leafy propagula or filamentous gemmae clustered in leaf axils. |
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Herzogiella striatella |
Hypnaceae |
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Phenology | Capsules mature summer. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Shaded soil and humus, acidic cliffs and rock, rotten logs, stumps, base of trees, exposed tree roots | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | low to high elevations (0-2000 m) (low to high elevations (0-6600 ft)) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
AK; CT; DE; GA; KY; MA; MD; ME; MI; MO; NC; NH; NJ; NY; OH; PA; RI; TN; VA; VT; WA; WI; WV; BC; NB; NL; NS; ON; PE; QC; Greenland; Europe; Asia
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Nearly worldwide; most diverse in subtropics and tropics |
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Discussion | Herzogiella striatella is recognized by its close, squarrose to squarrose-recurved leaves with 2–4 rows of abruptly inflated, hyaline or orange to red, decurrent cells extending 4–6 cells down the stem. This species has a disjunct distribution in North America, occurring commonly in the eastern third of the continent and more rarely in the western part in British Columbia, Alaska, and Washington. Herzogiella striatella may have a closer relationship to H. cylindricarpa (Cardot) Z. Iwatsuki of Mexico, Central America, South America, and Africa than to other North American species, as revealed by a study of spore ornamentation (R. R. Ireland 1990). The report for Alabama by Ireland (1991b) is an error. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 60, species ca. 600 (19 genera, 62 species in the flora). Hypnaceae are taxonomically problematic; the family once held a high proportion of pleurocarpous species, but as genera are monographed, they are often placed in other families. The distinctive hypnoid peristome has been used as a significant feature, but this is not entirely reliable. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 28, p. 528. | FNA vol. 28, p. 515. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Hypnaceae > Herzogiella | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Synonyms | Leskea striatella, Dolichotheca striatella, H. fitzgeraldii, H. muehlenbeckii, Isopterygium striatellum, Plagiothecium muehlenbeckii, P. striatellum, Sharpiella striatella | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | (Bridel) Z. Iwatsuki: J. Hattori Bot. Lab. 33: 374. (1970) | Schimper | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Web links |