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King's aster

Habit Perennials, densely cespitose, dwarf, 1–12 cm; caudices woody, branched, from stout taproots. Subshrubs or perennials, 30–60 cm cespitose; rhizomes woody, elongate, branched.
Stems

1–5+, loosely erect, densely long-stipitate-glandular.

1–20+, erect, branched, straight to ± flexuous, long- and short-stipitate-glandular, more densely so distally.

Leaves

mostly basal and cauline, basal and proximal cauline tufted, often marcescent; petiolate to sessile (distal), petioles 10–32 mm, bases sheathing;

blades oblanceolate to spatulate, 10–80(–120) × 15(–20) mm, progressively reduced distally, thick, firm, bases ± attenuate, margins entire or irregularly serrate, eciliate, teeth apiculate to ± spinulose, apices obtuse to acute, mucronate-indurate or callous, faces glabrous.

cauline, proximal reduced and deciduous by flowering, coriaceous, rigid;

sessile;

blades oblong to oblong-ovate or ovate to nearly orbiculate, 10–45 × 4–30 mm, gradually reduced distally, bases cordate-clasping, main veins marked, margins sharply spinulose-serrate, scabridulous, short-stipitate-glandular, apices acute to obtuse, mucronate, faces scabrellous, ± short-stipitate-glandular.

Peduncles

stipitate-glandular [bracts not observed].

0.3–4 cm, densely stipitate-glandular;

bracts 0–3, often proximal to heads and appearing to be part of involucres, foliaceous, margins entire or ± serrate.

Involucres

hemispheric-campanulate, 8–12 mm.

campanulate, 8.5–12 mm.

Ray florets

13–27;

corollas white to lavender, 8–17(–20) × 1.7–2.2(–3) mm.

17–20;

laminae purple, 15–22 × 1.2–2 mm.

Disc florets

29–47;

corollas pale yellow, 5.5–(6–8) mm, barely ampliate, tubes 2.6–2.9 mm, slightly longer than funnelform throats 1.9–2.3 mm, lobes erect to spreading, lanceolate, 0.8–1 mm.

ca. 29;

corollas yellow turning purplish, barely ampliate, ca. 8.2 mm, tubes shorter than funnelform throats (ca. 2.5 mm), lobes erect to spreading, triangular, 0.6–0.8 mm.

Phyllaries

in 4–5 series, often partly purplish (especially inner), ± keeled, lanceolate to linear-lanceolate, margins narrowly scarious, erose, sometimes stipitate-glandular (particularly on green parts), green zones 1/2–2/3 distal portions and full width (outer) to less than 1/6 and along midveins (inner), apices acute to long-acuminate, squarrose (outer and mid) to appressed (inner), faces glabrous, stipitate-glandular (green parts).

(ca. 40) in 5–6 series, spreading, sometimes purplish tinged (inner or bases of outer), keeled, linear-lanceolate to oblanceolate, lance-oblong, or lanceolate, coriaceous, margins narrowly scarious, hyaline, sometimes purplish distally (inner), erose, short-stipitate-glandular along foliaceous distal parts, apices acute and spinulose to long-cuminate (innermost), often purplish-margined, faces short-stipitate-glandular;

outer apically broadly foliaceous 1/3–9/10 (foliaceous part often wider than base), inner usually narrowly foliaceous 1/3 or less, sometimes green parts lanceolate apically along midnerves or none.

Heads

1–5, borne singly or in ± corymbiform arrays.

1–30+ in open corymbiform arrays or borne singly at ends of long, leafy branches.

Cypselae

brown, fusiform, slightly compressed, 2.5–3.5 mm, 8–10-nerved, faces ± densely strigillose;

pappi of yellowish bristles ca. 6 mm.

brown, fusiform, ± compressed [mature size not available], ribs 7–10, glabrous;

pappi of (ca. 40) tawny bristles 7–7.5 mm.

2n

= 18.

Herrickia kingii

Herrickia horrida

Phenology Flowering Jul–Sep.
Habitat Rocky hillsides, steep, narrow, brushy, canyon bottoms, with Quercus, Juniperus, Fallugia, and Forestiera
Elevation 1600–2900 m (5200–9500 ft)
Distribution
from FNA
UT
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; NM
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Herrickia kingii is known only from the Wasatch and Canyon mountains. The generic assignment of this species has been debated (e.g., A. Cronquist and D. D. Keck 1957; G. L. Nesom 1991e; Cronquist 1994). S. L. Welsh et al. (1987) underlined similarities with “Aster” alpigenus (= Oreostemma alpigena), among others. A recent molecular phylogenetic analysis (L. Brouillet et al. 2004) showed that H. kingii belongs to the Eurybia complex, possibly as a basal member of Herrickia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Herrickia horrida is known only from the Animas River basin, in Colfax County, northern New Mexico, and Las Animas County, southern Colorado.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves entire
var. kingii
1. Leaves (at least some basal and proximal) irregularly serrate
var. barnebyana
Source FNA vol. 20, p. 362. FNA vol. 20, p. 364.
Parent taxa Asteraceae > tribe Astereae > Herrickia Asteraceae > tribe Astereae > Herrickia
Sibling taxa
H. glauca, H. horrida, H. wasatchensis
H. glauca, H. kingii, H. wasatchensis
Subordinate taxa
H. kingii var. barnebyana, H. kingii var. kingii
Synonyms Aster kingii, Machaeranthera kingii, Tonestus kingii Aster horridus, Eurybia horrida
Name authority (D. C. Eaton) Brouillet: Sida 21: 898. (2004) Wooton & Standley: Contr. U.S. Natl. Herb. 16: 186, plate 50. (1913)
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