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Cleveland's tarweed, hayfield tarweed

hayfield tarweed, tarweed

Habit Annuals, 5–80 cm.
Stems

± erect.

Leaves

blades 50–175+ × 2–8(–12+) mm.

mostly cauline;

proximal opposite (forming winter–spring rosettes, sometimes present at flowering), mostly alternate;

sessile;

blades narrowly elliptic to linear or lance-linear, margins serrulate or entire, faces puberulent, hispidulous, hirsute, strigose, sericeous, or villous (distal leaves sometimes stipitate-glandular as well).

Involucres

hemispheric to ± urceolate or globose, 3.5–12 × 3–8+ mm.

hemispheric to ± urceolate or globose, 3–8+ mm diam. (usually subtended by calyculi of 5–7 bractlets in H. congesta subsp. calyculata).

Receptacles

flat to conic, glabrous, paleate (paleae connate, forming cells around all or most individual disc florets, scarious, ± deliquescent).

Ray florets

5–14, pistillate, fertile;

corollas white or yellow (often purple-veined abaxially).

Ray laminae

5–12 mm.

Disc florets/Disc corollas

2.5–3.5 mm.

5–60+, functionally staminate;

corollas white or yellow, tubes shorter than funnelform throats, lobes 5, deltate (anthers ± dark purple; styles glabrous proximal to branches).

Phyllaries

5–14 in 1 series (linear to lanceolate or oblanceolate, herbaceous, each usually 1/2 enveloping a subtended ray floret proximally, abaxially pubescent to hirsute or villous, and stipitate-glandular).

Heads

radiate, borne singly or in ± paniculiform, racemiform, or spiciform arrays or in glomerules.

Cypselae

black, 2–3.5 mm.

Disc cypselae

0;

pappi 0.

Ray cypselae

± obcompressed, abaxially gibbous, basal attachments oblique, apices sometimes beaked, beaks inconspicuous, straight, diameters greater than lengths, faces glabrous;

pappi 0.

Peduncular

bracts: pit-glands, tack-glands, and/or spines 0.

x

= 14.

Hemizonia congesta

Hemizonia

Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from USDA
w North America
[BONAP county map]
Discussion

Subspecies 6 (6 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 1.

Hemizonia as often circumscribed (to include Centromadia and Deinandra) is polyphyletic; molecular phylogenetic studies have indicated that Centromadia and Deinandra are more closely related to Calycadenia, Holocarpha, and Osmadenia than to Hemizonia in the strict sense, which is most closely related to Blepharizonia, also with 2n = 28 (S. Carlquist et al. 2003). Crosses between taxa of Hemizonia have yielded hybrids of minimal to full fertility; natural hybrids also have been documented (J. Clausen 1951). All taxa in Hemizonia are self-incompatible. The name H. congesta subsp. congesta has been sometimes misapplied to plants treated here as H. congesta subsp. lutescens; the type of H. congesta corresponds to plants that have been called H. leucocephala, a synonym of H. congesta subsp. congesta (B. G. Baldwin et al. 2001).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Heads in spiciform-racemiform to spiciform-paniculiform arrays (heads ± sessile); ray laminae white, abaxially purple-veined
→ 2
1. Heads in racemiform or paniculiform arrays or in glomerules; ray laminae white or yellow, abaxially purple-veined or not
→ 3
2. Peduncular bracts not or barely surpassing phyllaries; calyculi 0; phyllaries 3–7 mm, apices usually shorter than bodies; cypsela widths 0.6–0.75 times lengths
subsp. clevelandii
2. Peduncular bracts (and calyculi) often notably surpassing phyllaries; calyculi usually of 5–7, sometimes 0, bractlets; phyllaries 6–12 mm, apices usually longer than bodies; cypsela widths 0.5–0.6 times lengths
subsp. calyculata
3. Leaves usually puberulent and eglandular throughout, rarely the distal villous and glandular; heads in paniculiform arrays; ray laminae white, not purple-veined
subsp. tracyi
3. Leaves pubescent, sericeous, or villous, all or the distal glandular; heads in paniculiform arrays or in glomerules; ray laminae white or yellow, abaxially purple-veined
→ 4
4. Leaves ± villous (the distal with hairs at margins often longer); some or all heads usually in glomerules, sometimes in corymbiform or paniculiform arrays; phyllaries 6–10 mm, apices usually longer than bodies; ray laminae white
subsp. congesta
4. Leaves pubescent, villous, or sericeous (the distal with hairs at margins not notably longer); heads in paniculiform arrays; phyllaries 3–7(–8) mm, apices usually shorter than bodies; ray laminae white or yellow
→ 5
5. Ray laminae white
subsp. luzulifolia
5. Ray laminae yellow
subsp. lutescens
Source FNA vol. 21, p. 292. FNA vol. 21, p. 291. Authors: Bruce G. Baldwin, John L. Strother.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Madiinae > Hemizonia Asteraceae > tribe Heliantheae > subtribe Madiinae
Subordinate taxa
H. congesta subsp. calyculata, H. congesta subsp. clevelandii, H. congesta subsp. congesta, H. congesta subsp. lutescens, H. congesta subsp. luzulifolia, H. congesta subsp. tracyi
H. congesta
Name authority de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 5: 692. (1836) de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 5: 692. (1836)
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