Helianthus tuberosus |
Helianthus petiolaris |
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girasole, Jerusalem artichoke, Jerusalem sunflower, topinambour, tuberous sunflower |
prairie sunflower, unknown |
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Habit | Perennials, 50–200+ cm (rhizomatous, producing tubers late in growing season). | Annuals, 40–200 cm. | ||||
Stems | erect, scabro-hispid to hirsute (sometimes glaucous). |
erect, usually densely canescent, hispid, or strigillose, rarely ± hirsute or glabrate. |
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Leaves | mostly cauline; opposite or alternate proximally, usually alternate distally; petioles 2–8 cm (often ± winged); blades (3-nerved from near bases) lanceolate to ovate, 10–23 × 7–15 cm, bases broadly to narrowly cuneate, margins entire or serrate (flat), abaxial faces puberulent or hirsutulous to tomentulose and gland-dotted (abaxial) or ± scabrous (adaxial). |
mostly cauline; mostly alternate; petioles 2–4 cm; blades (often bluish green) lanceolate to deltate-ovate or ovate, 4–15 × 1–8 cm, bases subcordate or truncate to cuneate, margins entire or ± serrate, abaxial faces strigose, sparsely to densely, or not at all, gland-dotted. |
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Peduncles | 1–15 cm. |
4–15(–40) cm. |
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Involucres | hemispheric, (10–25 ×) 8–12 mm. |
± hemispheric, 10–24 mm diam. |
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Ray florets | 10–20; laminae 25–40 mm. |
10–30; laminae 15–20 mm. |
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Disc florets | 60+; corollas 6–7 mm, lobes yellow; anthers dark brown or black, appendages dark or yellowish. |
50–100+; corollas 4.5–6 mm, lobes usually reddish, rarely yellow; anthers reddish to purplish, appendages purplish (style branches reddish). |
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Phyllaries | (often dark green, drying nearly black) 22–35 (bases appressed, apices ± spreading, sometimes reflexed in fruit), lanceolate, 8.5–15 × 2–4 mm (subequal), (margins ciliate) apices acuminate, abaxial faces hispidulous or puberulent, gland-dotted. |
14–25, lance-linear to lanceolate to lance-ovate, 10–14 × 1–4(–5) mm, (margins sometimes ciliate) apices short-attenuate, abaxial faces usually hispidulous, rarely sparsely hirsute to glabrate. |
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Heads | 3–15. |
1–5. |
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Cypselae | 5–7 mm, glabrous or distally hairy; pappi of 2 aristate scales 1.9–3 mm plus 0–1 deltate scales 0.5–0.8 mm. |
3–4.5 mm, ± villous; pappi of 2 aristate scales 1.5–3 mm plus 0–2 erose scales 0.3–0.5 mm. |
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Paleae | 8–9 mm, 3-toothed (apices hairy). |
4.5–7.5 mm, 3-toothed, middle teeth ± ciliate or bearded, hairs whitish, 0.5–0.7 mm. |
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2n | = 102. |
= 34. |
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Helianthus tuberosus |
Helianthus petiolaris |
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Phenology | Flowering late summer–fall. | |||||
Habitat | Roadsides, fields, waste areas | |||||
Elevation | 0–1000(–1500) m (0–3300(–4900) ft) | |||||
Distribution |
AL; AR; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; MB; NB; NS; ON; PE; QC; SK; cultivated and adventive in Europe
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AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; LA; MA; ME; MI; MN; MO; MT; NC; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; SC; SD; TN; TX; UT; VA; WA; WI; WV; WY; AB; MB; ON; SK
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Discussion | Helianthus tuberosus is variable, probably in part stemming from hybridization with other polyploids, including H. pauciflorus, H. resinosus, and H. strumosus. Helianthus tuberosus is so widely spread as a weedy species that its original distribution is difficult to discern. It has been used as a food plant for its tubers by native Americans (although not necessarily domesticated or even cultivated); it has been developed as a crop primarily in Europe, where it has become widely naturalized. The common name Jerusalem artichoke is a misnomer, as explained by C. B. Heiser (1976). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). Helianthus petiolaris is adventive beyond western North America. A third subspecies has yet to be named at that rank in Helianthus petiolaris; it has been called H. petiolaris var. canescens A. Gray. It differs in having stems, leaves, and phyllaries densely canescent and abaxial faces of leaves densely gland-dotted. It is additionally characterized by peduncles usually ebracteate, phyllaries 1–2 mm wide, disc corolla throats gradually narrowed distal to slight, not densely hairy basal bulges, and 2n = 34. It flowers late spring through late summer and grows on sandy soils in open areas at (10–)1000–2300 m in Arizona, California, Nevada, New Mexico, and Texas and in Mexico. It was treated as H. niveus (Bentham) Brandegee subsp. canescens (A. Gray) Heiser by C. B. Heiser et al. (1969); molecular and morphologic data appear to favor a placement within H. petiolaris. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 21, p. 161. | FNA vol. 21, p. 152. | ||||
Parent taxa | Asteraceae > tribe Heliantheae > subtribe Helianthinae > Helianthus | Asteraceae > tribe Heliantheae > subtribe Helianthinae > Helianthus | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | H. tomentosus, H. tuberosus var. subcanescens | |||||
Name authority | Linnaeus: Sp. Pl. 2: 905. (1753) | Nuttall: J. Acad. Nat. Sci. Philadelphia 2: 115. (1821) | ||||
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