Helianthus tuberosus |
Helianthus nuttallii |
|||||||||
---|---|---|---|---|---|---|---|---|---|---|
girasole, Jerusalem artichoke, Jerusalem sunflower, topinambour, tuberous sunflower |
hélianthe de Nuttall, Nuttall's sunflower |
|||||||||
Habit | Perennials, 50–200+ cm (rhizomatous, producing tubers late in growing season). | Perennials, 100–400 cm (rhizomatous). | ||||||||
Stems | erect, scabro-hispid to hirsute (sometimes glaucous). |
(usually yellow-brown or greenish, sometimes glaucous) erect, glabrous, hispid, ± hirsute, or scabrous. |
||||||||
Leaves | mostly cauline; opposite or alternate proximally, usually alternate distally; petioles 2–8 cm (often ± winged); blades (3-nerved from near bases) lanceolate to ovate, 10–23 × 7–15 cm, bases broadly to narrowly cuneate, margins entire or serrate (flat), abaxial faces puberulent or hirsutulous to tomentulose and gland-dotted (abaxial) or ± scabrous (adaxial). |
cauline; all or mostly opposite to mostly alternate; petioles 0.5–1.5 cm; blades light to dark green, 3-nerved (distal to bases) ± lanceolate to ± ovate, 4–20 × 0.8–4 cm, bases cuneate, margins entire or ± serrate (flat), abaxial faces hispid to hispidulous or tomentulose to villous-tomentose, gland-dotted. |
||||||||
Peduncles | 1–15 cm. |
1–18 cm. |
||||||||
Involucres | hemispheric, (10–25 ×) 8–12 mm. |
hemispheric, 10–20 mm diam. |
||||||||
Ray florets | 10–20; laminae 25–40 mm. |
10–21; laminae 20–25 mm (abaxial faces not gland-dotted). |
||||||||
Disc florets | 60+; corollas 6–7 mm, lobes yellow; anthers dark brown or black, appendages dark or yellowish. |
60+; corollas 5–7 mm, lobes yellow; anthers dark brown to black, appendages yellow. |
||||||||
Phyllaries | (often dark green, drying nearly black) 22–35 (bases appressed, apices ± spreading, sometimes reflexed in fruit), lanceolate, 8.5–15 × 2–4 mm (subequal), (margins ciliate) apices acuminate, abaxial faces hispidulous or puberulent, gland-dotted. |
30–38 (loose, spreading), lanceolate to lance-ovate, 8–16 × 1.5–3 mm, (margins ciliate) apices acute to acuminate, abaxial faces usually ± strigose, rarely glabrate, not gland-dotted. |
||||||||
Heads | 3–15. |
1–6. |
||||||||
Cypselae | 5–7 mm, glabrous or distally hairy; pappi of 2 aristate scales 1.9–3 mm plus 0–1 deltate scales 0.5–0.8 mm. |
3–5 mm, glabrate; pappi of 2 aristate scales 2.2–4.5 mm plus 0–2 lanceolate scales 0.5–1 mm. |
||||||||
Paleae | 8–9 mm, 3-toothed (apices hairy). |
8–12 mm, 3-toothed. |
||||||||
2n | = 102. |
= 34 (subspecies unknown). |
||||||||
Helianthus tuberosus |
Helianthus nuttallii |
|||||||||
Phenology | Flowering late summer–fall. | |||||||||
Habitat | Roadsides, fields, waste areas | |||||||||
Elevation | 0–1000(–1500) m (0–3300(–4900) ft) | |||||||||
Distribution |
AL; AR; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; MB; NB; NS; ON; PE; QC; SK; cultivated and adventive in Europe
|
AZ; CA; CO; IA; ID; KS; MN; MO; MT; ND; NE; NM; NV; OK; OR; SD; UT; WA; WY; AB; BC; MB; ON; QC; SK
|
||||||||
Discussion | Helianthus tuberosus is variable, probably in part stemming from hybridization with other polyploids, including H. pauciflorus, H. resinosus, and H. strumosus. Helianthus tuberosus is so widely spread as a weedy species that its original distribution is difficult to discern. It has been used as a food plant for its tubers by native Americans (although not necessarily domesticated or even cultivated); it has been developed as a crop primarily in Europe, where it has become widely naturalized. The common name Jerusalem artichoke is a misnomer, as explained by C. B. Heiser (1976). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 3 (3 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||||||
Key |
|
|||||||||
Source | FNA vol. 21, p. 161. | FNA vol. 21, p. 166. | ||||||||
Parent taxa | Asteraceae > tribe Heliantheae > subtribe Helianthinae > Helianthus | Asteraceae > tribe Heliantheae > subtribe Helianthinae > Helianthus | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | H. tomentosus, H. tuberosus var. subcanescens | |||||||||
Name authority | Linnaeus: Sp. Pl. 2: 905. (1753) | Torrey & A. Gray: Fl. N. Amer. 2: 324. (1842) | ||||||||
Web links |
|
|