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girasole, Jerusalem artichoke, Jerusalem sunflower, topinambour, tuberous sunflower

ashy sunflower, downy sunflower

Habit Perennials, 50–200+ cm (rhizomatous, producing tubers late in growing season). Perennials, 50–150+ cm (rhizoma-tous).
Stems

erect, scabro-hispid to hirsute (sometimes glaucous).

erect, hirsute to villous.

Leaves

mostly cauline; opposite or alternate proximally, usually alternate distally;

petioles 2–8 cm (often ± winged);

blades (3-nerved from near bases) lanceolate to ovate, 10–23 × 7–15 cm, bases broadly to narrowly cuneate, margins entire or serrate (flat), abaxial faces puberulent or hirsutulous to tomentulose and gland-dotted (abaxial) or ± scabrous (adaxial).

mostly cauline; mostly opposite (sometimes alternate among heads);

sessile;

blades (ashy or gray-green, 3-nerved distal to bases) lanceolate to broadly ovate, 5.5–14.5 × 1.8–6.5 cm, bases rounded to cordate, margins entire or serrulate, abaxial faces hispid to tomentose, gland-dotted.

Peduncles

1–15 cm.

0.1–15 cm.

Involucres

hemispheric, (10–25 ×) 8–12 mm.

broadly hemispheric, 12–25 mm diam.

Ray florets

10–20;

laminae 25–40 mm.

17–22;

laminae 25–30 mm (abaxial faces densely gland-dotted).

Disc florets

60+;

corollas 6–7 mm, lobes yellow;

anthers dark brown or black, appendages dark or yellowish.

75+;

corollas 6–7.5 mm, lobes yellow;

anthers dark, appendages dark.

Phyllaries

(often dark green, drying nearly black) 22–35 (bases appressed, apices ± spreading, sometimes reflexed in fruit), lanceolate, 8.5–15 × 2–4 mm (subequal), (margins ciliate) apices acuminate, abaxial faces hispidulous or puberulent, gland-dotted.

30–40, lanceolate, (5–)10–16 × 2–3.5 mm, apices usually acute, sometimes acuminate, abaxial faces densely hispid to villous or tomentose, densely gland-dotted.

Heads

3–15.

1–15.

Cypselae

5–7 mm, glabrous or distally hairy;

pappi of 2 aristate scales 1.9–3 mm plus 0–1 deltate scales 0.5–0.8 mm.

3.5–4 mm, distally villous;

pappi of 2 aristate scales 2.8–3.2 mm.

Paleae

8–9 mm, 3-toothed (apices hairy).

(oblanceolate) 9–11 mm, entire (1-toothed, densely hairy, densely gland-dotted).

2n

= 102.

= 34.

Helianthus tuberosus

Helianthus mollis

Phenology Flowering late summer–fall. Flowering summer–early fall.
Habitat Roadsides, fields, waste areas Prairies, roadsides
Elevation 0–1000(–1500) m (0–3300(–4900) ft) 10–600+ m (0–2000+ ft)
Distribution
from FNA
AL; AR; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; MB; NB; NS; ON; PE; QC; SK; cultivated and adventive in Europe
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MS; NC; NE; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WI; WV; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

Helianthus tuberosus is variable, probably in part stemming from hybridization with other polyploids, including H. pauciflorus, H. resinosus, and H. strumosus. Helianthus tuberosus is so widely spread as a weedy species that its original distribution is difficult to discern. It has been used as a food plant for its tubers by native Americans (although not necessarily domesticated or even cultivated); it has been developed as a crop primarily in Europe, where it has become widely naturalized. The common name Jerusalem artichoke is a misnomer, as explained by C. B. Heiser (1976).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Helianthus mollis is introduced in Ontario and adventive in the eastern United States (e.g., Maine), where it is continuing to spread, particularly along roads. Natural hybrids between H. mollis and H. occidentalis have been named H. cinereus Torrey & A. Gray (R. C. Jackson and A. T. Guard 1957); they differ from H. mollis by having smaller heads with fewer ray florets and narrower leaves with cuneate bases. Hybrids of H. mollis with H. giganteus have been called H. doronicoides Lamarck (Jackson 1956).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 21, p. 161. FNA vol. 21, p. 163.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Helianthinae > Helianthus Asteraceae > tribe Heliantheae > subtribe Helianthinae > Helianthus
Sibling taxa
H. agrestis, H. angustifolius, H. annuus, H. anomalus, H. argophyllus, H. arizonensis, H. atrorubens, H. bolanderi, H. californicus, H. carnosus, H. ciliaris, H. cusickii, H. debilis, H. decapetalus, H. deserticola, H. divaricatus, H. eggertii, H. exilis, H. floridanus, H. giganteus, H. glaucophyllus, H. gracilentus, H. grosseserratus, H. heterophyllus, H. hirsutus, H. laciniatus, H. laevigatus, H. longifolius, H. maximiliani, H. microcephalus, H. mollis, H. neglectus, H. niveus, H. nuttallii, H. occidentalis, H. paradoxus, H. pauciflorus, H. petiolaris, H. porteri, H. praecox, H. pumilus, H. radula, H. resinosus, H. salicifolius, H. schweinitzii, H. silphioides, H. simulans, H. smithii, H. strumosus, H. verticillatus, H. ×laetiflorus
H. agrestis, H. angustifolius, H. annuus, H. anomalus, H. argophyllus, H. arizonensis, H. atrorubens, H. bolanderi, H. californicus, H. carnosus, H. ciliaris, H. cusickii, H. debilis, H. decapetalus, H. deserticola, H. divaricatus, H. eggertii, H. exilis, H. floridanus, H. giganteus, H. glaucophyllus, H. gracilentus, H. grosseserratus, H. heterophyllus, H. hirsutus, H. laciniatus, H. laevigatus, H. longifolius, H. maximiliani, H. microcephalus, H. neglectus, H. niveus, H. nuttallii, H. occidentalis, H. paradoxus, H. pauciflorus, H. petiolaris, H. porteri, H. praecox, H. pumilus, H. radula, H. resinosus, H. salicifolius, H. schweinitzii, H. silphioides, H. simulans, H. smithii, H. strumosus, H. tuberosus, H. verticillatus, H. ×laetiflorus
Synonyms H. tomentosus, H. tuberosus var. subcanescens H. mollis var. cordatus
Name authority Linnaeus: Sp. Pl. 2: 905. (1753) Lamarck: in J. Lamarck et al., Encycl. 3: 85. (1789)
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