Helianthus tuberosus |
Helianthus exilis |
|
|---|---|---|
|
girasole, hélianthe scrofuleux, Jerusalem artichoke, Jerusalem sunflower, rough sunflower, topinambour, tuberous sunflower |
Eggert's sunflower, serpentine sunflower |
|
| Habit | Perennials, 50–200+ cm (rhizomatous, producing tubers late in growing season). | Annuals, 30–100 cm. |
| Stems | erect, scabro-hispid to hirsute (sometimes glaucous). |
(often reddish) erect, hairy. |
| Leaves | mostly cauline; opposite or alternate proximally, usually alternate distally; petioles 2–8 cm (often ± winged); blades (3-nerved from near bases) lanceolate to ovate, 10–23 × 7–15 cm, bases broadly to narrowly cuneate, margins entire or serrate (flat), abaxial faces puberulent or hirsutulous to tomentulose and gland-dotted (abaxial) or ± scabrous (adaxial). |
mostly cauline; mostly alternate; petioles 0.7–2.5 cm; blades lance-linear to lance-ovate or ovate, 3–15 × 0.5–3 cm, bases cuneate, margins usually entire or shallowly serrate, abaxial faces gland-dotted. |
| Peduncles | 1–15 cm. |
(1–)3–13(–20) cm. |
| Involucres | hemispheric, (10–25 ×) 8–12 mm. |
hemispheric, 15–20 mm diam. |
| Ray florets | 10–20; laminae 25–40 mm. |
10–13; laminae 14–20 mm. |
| Disc florets | 60+; corollas 6–7 mm, lobes yellow; anthers dark brown or black, appendages dark or yellowish. |
50+; corollas 4–6 mm, lobes reddish; anthers reddish purple, appendages usually purplish (style branches reddish or yellow). |
| Phyllaries | (often dark green, drying nearly black) 22–35 (bases appressed, apices ± spreading, sometimes reflexed in fruit), lanceolate, 8.5–15 × 2–4 mm (subequal), (margins ciliate) apices acuminate, abaxial faces hispidulous or puberulent, gland-dotted. |
12–17, lanceolate, 8–17 × 3–4 mm, apices narrowed gradually, abaxial faces hirsute. |
| Heads | 3–15. |
1–7. |
| Cypselae | 5–7 mm, glabrous or distally hairy; pappi of 2 aristate scales 1.9–3 mm plus 0–1 deltate scales 0.5–0.8 mm. |
(2.5–)3–3.5(–4) mm, glabrate; pappi of 2 lanceolate scales 1.7–2.7 mm. |
| Paleae | 8–9 mm, 3-toothed (apices hairy). |
9.5–10 mm, 3-toothed (middle teeth surpassing discs, apices greenish or yellow-brown, glabrous). |
| 2n | = 102. |
= 34. |
Helianthus tuberosus |
Helianthus exilis |
|
| Phenology | Flowering late summer–fall. | Flowering summer–early fall. |
| Habitat | Roadsides, fields, waste areas | Gravelly streamsides on serpentine |
| Elevation | 0–1000(–1500) m [0–3300(–4900) ft] | 100–1400 m [300–4600 ft] |
| Distribution |
AL; AR; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; MB; NB; NS; ON; PE; QC; SK; cultivated and adventive in Europe
|
CA |
| Discussion | Helianthus tuberosus is variable, probably in part stemming from hybridization with other polyploids, including H. pauciflorus, H. resinosus, and H. strumosus. Helianthus tuberosus is so widely spread as a weedy species that its original distribution is difficult to discern. It has been used as a food plant for its tubers by native Americans (although not necessarily domesticated or even cultivated); it has been developed as a crop primarily in Europe, where it has become widely naturalized. The common name Jerusalem artichoke is a misnomer, as explained by C. B. Heiser (1976). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Helianthus exilis is very similar morphologically to H. bolanderi and is often included in it; there are discrete differences between the two for fatty acid composition (C. E. Rogers et al. 1982) as well as some molecular markers (L. H. Rieseberg et al. 1988). It is listed as of concern by the California Native Plant Society. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| Parent taxa | ||
| Sibling taxa | ||
| Synonyms | H. tomentosus, H. tuberosus var. subcanescens | |
| Name authority | Linnaeus: Sp. Pl. 2: 905. (1753) | A. Gray: Proc. Amer. Acad. Arts 6: 545. (1865) |
| Source | FNA vol. 21, p. 161. | FNA vol. 21, p. 150. |
| Web links |
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