The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

hélianthe de maximilien, Maximilian sunflower, Maximilian's sunflower

Habit Perennials, 50–300 cm (rhizomatous). Annuals, perennials, subshrubs, or shrubs, (2–)20–500 cm.
Stems

erect, 5–30 dm, scabrous to scabro-hispidulous.

Leaves

cauline; mostly alternate;

petioles 0–2 cm;

blades (light green to gray-green, 1-nerved, conduplicate) lanceolate, 10–30 × 2–5.5 cm, bases cuneate, margins usually entire, sometimes serrulate, abaxial faces scabrous to scabro-hispid, gland-dotted.

usually mostly cauline, rarely mostly basal; mostly opposite (distal often alternate) or mostly alternate, rarely whorled; petiolate or sessile;

blades (often 3- or 5-nerved) orbiculate or deltate to lanceolate or linear (and intermediate shapes), sometimes lobed, ultimate margins entire or toothed, faces usually hairy (often hispid to scabrellous), often gland-dotted.

Peduncles

1–11 cm.

Involucres

hemispheric, 13–28 mm diam.

cylindric to hemispheric or rotate.

Receptacles

usually flat or convex, sometimes hemispheric or conic, usually paleate (epaleate in Lagascea; paleae usually falling, sometimes persistent, mostly oblong to linear, often conduplicate, herbaceous to scarious, apices sometimes ± 3-lobed or -toothed; becoming hardened perigynia completely investing cypselae in Sclerocarpus).

Ray florets

10–25;

laminae (15–)25–40 mm.

0 or 5–30(–100+), neuter, or styliferous and sterile;

corollas usually yellow to orange, sometimes partly or wholly brown to purple or red-brown to reddish.

Disc florets

75+;

corollas 5–7 mm, lobes yellow;

anthers dark brown or black;

appendages usually yellow, sometimes partly dark.

1 (Lagascea) or 10–200(–1000+), bisexual, fertile;

corollas usually yellow to orange, sometimes brown to purple or reddish to red-brown, tubes shorter than or about equaling cylindric or funnelform throats, lobes 5, ± deltate (equal);

anther thecae dark or pale;

stigmatic papillae usually continuous.

Phyllaries

30–40, lanceolate, 14–20 × 2–3 mm, (margins ciliate) apices acute to attenuate, abaxial faces canescent, gland-dotted.

persistent, 4–45(–100+) in 1–6+ series (usually distinct, ± connate in Lagascea, broadly ovate to linear, subequal or unequal).

Calyculi

0.

Heads

(1–)3–15 (often in racemiform to spiciform arrays).

usually radiate, sometimes discoid, borne singly or in corymbiform (paniculiform or racemiform) arrays (glomerules or second-order heads of 1–2-flowered heads in Lagascea).

Cypselae

3–4 mm, glabrate;

pappi of 2 aristate scales 3–4.1 mm.

often ± compressed, flattened, biconvex, clavate, columnar, obovoid, obpyramidal, or prismatic (lengths usually 1–2 times diams.), glabrous or hairy (faces and/or angles);

pappi 0, or (often readily falling or fragile, sometimes persistent) usually of (1–)2(–8+) (sometimes aristate) scales and/or awns, sometimes coroniform.

Paleae

7–11 mm, entire or 3-toothed (apices greenish, mucronate, hairy).

2n

= 34.

Helianthus maximiliani

Asteraceae tribe Heliantheae subtribe Helianthinae

Phenology Flowering late summer–fall.
Habitat Prairies, fields, waste areas
Elevation 0–300(–2100+) m (0–1000(–6900+) ft)
Distribution
from FNA
AL; AR; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MS; NC; ND; NE; NJ; NY; OH; OK; PA; SC; SD; TN; TX; UT; VA; WA; WI; WV; WY; AB; BC; MB; ON; QC; SK; Mexico
[WildflowerSearch map]
[BONAP county map]
Mostly subtropical; tropical; and warm-temperate New World
Discussion

Helianthus maximiliani is introduced in eastern Ontario and in Quebec. It appears to be native to midcontinental prairie regions and has spread along railroads and highways into all areas of North America. Its wide dispersal may be aided by cultivation for its attractive, showy floral displays. In addition to the usually conduplicate, single-nerved leaves and spikelike arrangement of the heads, it is distinguished by the whitish-canescent indument of the leaves and stems and the long-attenuate phyllaries.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 17, species 363 (8 genera, 69 species, including 1 hybrid, in the flora).

Circumscription of Helianthinae adopted here (i.e., that of H. Robinson 1981) is narrower than traditional circumscriptions (e.g., T. F. Stuessy 1977[1978]), which included genera here treated in Ecliptinae, Galinsoginae, and Rudbeckiinae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Phyllaries 4–5 in ± 1 series (connate); disc florets 1(–2; the 1–2-flowered heads aggregated in glomerules or second-order heads)
Lagascea
1. Phyllaries 5–45(–100+) in (1–)2–6+ series (distinct); disc florets (10–)15–200(–1000+)
→ 2
2. Receptacles paleate (each palea completely investing and falling with a cypsela, eachforming a hardened perigynium)
Sclerocarpus
2. Receptacles paleate (paleae sometimes conduplicate, ± enfolding cypselae, not forming perigynia)
→ 3
3. Heads borne singly (peduncles usually distally dilated, fistulose)
Tithonia
3. Heads borne singly or in corymbiform, paniculiform, racemiform, or thyrsiform arrays (peduncles rarely, if ever, notably dilated or fistulose)
→ 4
4. Cypselae flattened, thin-margined
Simsia
4. Cypselae ± compressed, biconvex, or 3- or 4-angled, often obpyramidal
→ 5
5. Cypselae glabrous or glabrate; pappi 0
→ 6
5. Cypselae usually ± strigose, sometimes glabrous or glabrate; pappi of 2–6+, persistent, readily falling, or tardily falling scales
→ 8
6. Shrubs (leaves often lobed, lobes usually 3–9, ± linear)
Viguiera
6. Annuals or perennials (leaves not lobed)
→ 7
7. Annuals; leaf blades lanceolate to linear; involucres 5–6 mm diam.;phyllaries 11–17
Helianthus
7. Annuals or perennials; leaf blades lance-linear, lanceolate, ovate, rhombic, or rhombic-ovate; involucres 6–14 mm diam.; phyllaries14–25
Heliomeris
8. Shrubs; involucres 5–9 mm diam
Bahiopsis
8. Annuals or perennials; involucres (5–)7–40+ mm diam
→ 9
9. Pappi readily falling
Helianthus
9. Pappi persistent or tardily falling
Viguiera
Source FNA vol. 21, p. 166. FNA vol. 21, p. 135.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Helianthinae > Helianthus Asteraceae > tribe Heliantheae
Sibling taxa
H. agrestis, H. angustifolius, H. annuus, H. anomalus, H. argophyllus, H. arizonensis, H. atrorubens, H. bolanderi, H. californicus, H. carnosus, H. ciliaris, H. cusickii, H. debilis, H. decapetalus, H. deserticola, H. divaricatus, H. eggertii, H. exilis, H. floridanus, H. giganteus, H. glaucophyllus, H. gracilentus, H. grosseserratus, H. heterophyllus, H. hirsutus, H. laciniatus, H. laevigatus, H. longifolius, H. microcephalus, H. mollis, H. neglectus, H. niveus, H. nuttallii, H. occidentalis, H. paradoxus, H. pauciflorus, H. petiolaris, H. porteri, H. praecox, H. pumilus, H. radula, H. resinosus, H. salicifolius, H. schweinitzii, H. silphioides, H. simulans, H. smithii, H. strumosus, H. tuberosus, H. verticillatus, H. ×laetiflorus
Subordinate taxa
Bahiopsis, Helianthus, Heliomeris, Lagascea, Sclerocarpus, Simsia, Tithonia, Viguiera
Synonyms H. dalyi subtribe Lagasceinae
Name authority Schrader: Index Seminum (Göttingen) 1834: unpaged. (1835) Cassini ex Dumortier: Fl. Belg., 71. (1827)
Web links