Helianthus maximiliani |
Asteraceae tribe Heliantheae subtribe Helianthinae |
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hélianthe de maximilien, Maximilian sunflower, Maximilian's sunflower |
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Habit | Perennials, 50–300 cm (rhizomatous). | Annuals, perennials, subshrubs, or shrubs, (2–)20–500 cm. | ||||||||||||||||||||||||||||||||||||
Stems | erect, 5–30 dm, scabrous to scabro-hispidulous. |
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Leaves | cauline; mostly alternate; petioles 0–2 cm; blades (light green to gray-green, 1-nerved, conduplicate) lanceolate, 10–30 × 2–5.5 cm, bases cuneate, margins usually entire, sometimes serrulate, abaxial faces scabrous to scabro-hispid, gland-dotted. |
usually mostly cauline, rarely mostly basal; mostly opposite (distal often alternate) or mostly alternate, rarely whorled; petiolate or sessile; blades (often 3- or 5-nerved) orbiculate or deltate to lanceolate or linear (and intermediate shapes), sometimes lobed, ultimate margins entire or toothed, faces usually hairy (often hispid to scabrellous), often gland-dotted. |
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Peduncles | 1–11 cm. |
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Involucres | hemispheric, 13–28 mm diam. |
cylindric to hemispheric or rotate. |
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Receptacles | usually flat or convex, sometimes hemispheric or conic, usually paleate (epaleate in Lagascea; paleae usually falling, sometimes persistent, mostly oblong to linear, often conduplicate, herbaceous to scarious, apices sometimes ± 3-lobed or -toothed; becoming hardened perigynia completely investing cypselae in Sclerocarpus). |
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Ray florets | 10–25; laminae (15–)25–40 mm. |
0 or 5–30(–100+), neuter, or styliferous and sterile; corollas usually yellow to orange, sometimes partly or wholly brown to purple or red-brown to reddish. |
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Disc florets | 75+; corollas 5–7 mm, lobes yellow; anthers dark brown or black; appendages usually yellow, sometimes partly dark. |
1 (Lagascea) or 10–200(–1000+), bisexual, fertile; corollas usually yellow to orange, sometimes brown to purple or reddish to red-brown, tubes shorter than or about equaling cylindric or funnelform throats, lobes 5, ± deltate (equal); anther thecae dark or pale; stigmatic papillae usually continuous. |
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Phyllaries | 30–40, lanceolate, 14–20 × 2–3 mm, (margins ciliate) apices acute to attenuate, abaxial faces canescent, gland-dotted. |
persistent, 4–45(–100+) in 1–6+ series (usually distinct, ± connate in Lagascea, broadly ovate to linear, subequal or unequal). |
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Calyculi | 0. |
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Heads | (1–)3–15 (often in racemiform to spiciform arrays). |
usually radiate, sometimes discoid, borne singly or in corymbiform (paniculiform or racemiform) arrays (glomerules or second-order heads of 1–2-flowered heads in Lagascea). |
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Cypselae | 3–4 mm, glabrate; pappi of 2 aristate scales 3–4.1 mm. |
often ± compressed, flattened, biconvex, clavate, columnar, obovoid, obpyramidal, or prismatic (lengths usually 1–2 times diams.), glabrous or hairy (faces and/or angles); pappi 0, or (often readily falling or fragile, sometimes persistent) usually of (1–)2(–8+) (sometimes aristate) scales and/or awns, sometimes coroniform. |
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Paleae | 7–11 mm, entire or 3-toothed (apices greenish, mucronate, hairy). |
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2n | = 34. |
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Helianthus maximiliani |
Asteraceae tribe Heliantheae subtribe Helianthinae |
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Phenology | Flowering late summer–fall. | |||||||||||||||||||||||||||||||||||||
Habitat | Prairies, fields, waste areas | |||||||||||||||||||||||||||||||||||||
Elevation | 0–300(–2100+) m (0–1000(–6900+) ft) | |||||||||||||||||||||||||||||||||||||
Distribution |
AL; AR; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MS; NC; ND; NE; NJ; NY; OH; OK; PA; SC; SD; TN; TX; UT; VA; WA; WI; WV; WY; AB; BC; MB; ON; QC; SK; Mexico
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Mostly subtropical; tropical; and warm-temperate New World |
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Discussion | Helianthus maximiliani is introduced in eastern Ontario and in Quebec. It appears to be native to midcontinental prairie regions and has spread along railroads and highways into all areas of North America. Its wide dispersal may be aided by cultivation for its attractive, showy floral displays. In addition to the usually conduplicate, single-nerved leaves and spikelike arrangement of the heads, it is distinguished by the whitish-canescent indument of the leaves and stems and the long-attenuate phyllaries. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 17, species 363 (8 genera, 69 species, including 1 hybrid, in the flora). Circumscription of Helianthinae adopted here (i.e., that of H. Robinson 1981) is narrower than traditional circumscriptions (e.g., T. F. Stuessy 1977[1978]), which included genera here treated in Ecliptinae, Galinsoginae, and Rudbeckiinae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 21, p. 166. | FNA vol. 21, p. 135. | ||||||||||||||||||||||||||||||||||||
Parent taxa | Asteraceae > tribe Heliantheae > subtribe Helianthinae > Helianthus | Asteraceae > tribe Heliantheae | ||||||||||||||||||||||||||||||||||||
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Synonyms | H. dalyi | subtribe Lagasceinae | ||||||||||||||||||||||||||||||||||||
Name authority | Schrader: Index Seminum (Göttingen) 1834: unpaged. (1835) | Cassini ex Dumortier: Fl. Belg., 71. (1827) | ||||||||||||||||||||||||||||||||||||
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