Helianthus hirsutus |
Helianthus petiolaris |
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hairy sunflower |
prairie sunflower, unknown |
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Habit | Perennials, 100–200 cm (rhizomatous). | Annuals, 40–200 cm. | ||||
Stems | erect, hirsute. |
erect, usually densely canescent, hispid, or strigillose, rarely ± hirsute or glabrate. |
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Leaves | cauline; mostly opposite; petioles 0.4–2 cm; blades (3-nerved from bases) lanceolate to ovate, 6.5–18 × 1–8 cm, bases truncate to broadly rounded or cuneate, margins subentire to serrate (flat), abaxial faces ± hirsute, gland-dotted (adaxial not gland-dotted). |
mostly cauline; mostly alternate; petioles 2–4 cm; blades (often bluish green) lanceolate to deltate-ovate or ovate, 4–15 × 1–8 cm, bases subcordate or truncate to cuneate, margins entire or ± serrate, abaxial faces strigose, sparsely to densely, or not at all, gland-dotted. |
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Peduncles | 1–5 cm. |
4–15(–40) cm. |
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Involucres | hemispheric, 10–25 mm diam. |
± hemispheric, 10–24 mm diam. |
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Ray florets | 10–15; laminae 15–20 mm. |
10–30; laminae 15–20 mm. |
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Disc florets | 40+; corollas 5.5–6.5 mm, lobes yellow; anthers dark brown or black, appendages dark or yellowish. |
50–100+; corollas 4.5–6 mm, lobes usually reddish, rarely yellow; anthers reddish to purplish, appendages purplish (style branches reddish). |
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Phyllaries | 18–25 (usually loose, spreading, not reflexed), lanceolate, 7–12 × 2.5–3.5 mm, (margins ciliate) apices acute to short-acuminate, abaxial faces not gland-dotted. |
14–25, lance-linear to lanceolate to lance-ovate, 10–14 × 1–4(–5) mm, (margins sometimes ciliate) apices short-attenuate, abaxial faces usually hispidulous, rarely sparsely hirsute to glabrate. |
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Heads | 1–7. |
1–5. |
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Cypselae | 4–4.5 mm, glabrate or distally puberulent; pappi of 2 aristate scales 2.5–3.2 mm. |
3–4.5 mm, ± villous; pappi of 2 aristate scales 1.5–3 mm plus 0–2 erose scales 0.3–0.5 mm. |
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Paleae | 7–10 mm, 3-toothed (apices yellowish, hairy). |
4.5–7.5 mm, 3-toothed, middle teeth ± ciliate or bearded, hairs whitish, 0.5–0.7 mm. |
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2n | = 68. |
= 34. |
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Helianthus hirsutus |
Helianthus petiolaris |
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Phenology | Flowering late summer–fall. | |||||
Habitat | Dry, open sites, woodland edges, roadsides | |||||
Elevation | 10–900+ m (0–3000+ ft) | |||||
Distribution |
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; NC; NE; NY; OH; OK; PA; SC; TN; TX; VA; WI; WV; ON; Mexico (Coahuila, Nuevo León)
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AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; LA; MA; ME; MI; MN; MO; MT; NC; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; SC; SD; TN; TX; UT; VA; WA; WI; WV; WY; AB; MB; ON; SK
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Discussion | Helianthus hirsutus is distinguished from H. strumosus by hairy stems and usually yellow (as opposed to dark) anther appendages, and from H. divaricatus by petioles and leaf blades 3-nerved distal to bases. Mexican plants of H. hirsutus are sometimes labeled with the synonymous H. leptocaulis (S. Watson) S. F. Blake, and plants from Mexico and the southwestern United States often have leaf bases cuneate rather than truncate. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). Helianthus petiolaris is adventive beyond western North America. A third subspecies has yet to be named at that rank in Helianthus petiolaris; it has been called H. petiolaris var. canescens A. Gray. It differs in having stems, leaves, and phyllaries densely canescent and abaxial faces of leaves densely gland-dotted. It is additionally characterized by peduncles usually ebracteate, phyllaries 1–2 mm wide, disc corolla throats gradually narrowed distal to slight, not densely hairy basal bulges, and 2n = 34. It flowers late spring through late summer and grows on sandy soils in open areas at (10–)1000–2300 m in Arizona, California, Nevada, New Mexico, and Texas and in Mexico. It was treated as H. niveus (Bentham) Brandegee subsp. canescens (A. Gray) Heiser by C. B. Heiser et al. (1969); molecular and morphologic data appear to favor a placement within H. petiolaris. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 21, p. 157. | FNA vol. 21, p. 152. | ||||
Parent taxa | Asteraceae > tribe Heliantheae > subtribe Helianthinae > Helianthus | Asteraceae > tribe Heliantheae > subtribe Helianthinae > Helianthus | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | H. hirsutus var. stenophyllus, H. hirsutus var. trachyphyllus, H. stenophyllus | |||||
Name authority | Rafinesque: Ann. Nat. 1: 14. (1820) | Nuttall: J. Acad. Nat. Sci. Philadelphia 2: 115. (1821) | ||||
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