Helianthus heterophyllus |
Helianthus nuttallii |
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variableleaf sunflower, wetland sunflower |
hélianthe de Nuttall, Nuttall's sunflower |
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Habit | Perennials, 50–120 cm (with crown buds). | Perennials, 100–400 cm (rhizomatous). | ||||||||
Stems | erect, usually hispid to ± hirsute. |
(usually yellow-brown or greenish, sometimes glaucous) erect, glabrous, hispid, ± hirsute, or scabrous. |
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Leaves | mostly basal; mostly opposite; petioles 0–3 cm (broadly winged); blades (3-nerved distal to bases) ovate or lanceolate to spatulate, 6–28 × 1.2–4.3 cm, bases cuneate, margins entire (often revolute), abaxial faces hispid to ± hirsute, not gland-dotted (cauline leaves relatively few, narrowly lanceolate to linear, much smaller). |
cauline; all or mostly opposite to mostly alternate; petioles 0.5–1.5 cm; blades light to dark green, 3-nerved (distal to bases) ± lanceolate to ± ovate, 4–20 × 0.8–4 cm, bases cuneate, margins entire or ± serrate (flat), abaxial faces hispid to hispidulous or tomentulose to villous-tomentose, gland-dotted. |
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Peduncles | 10–15 cm. |
1–18 cm. |
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Involucres | broadly hemispheric, 15–25 mm diam. |
hemispheric, 10–20 mm diam. |
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Ray florets | 12–18; laminae 14–36 mm (abaxial faces not gland-dotted). |
10–21; laminae 20–25 mm (abaxial faces not gland-dotted). |
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Disc florets | 100+; corollas 5.5–6.5 mm, lobes reddish; anthers purplish, apendages purplish (style branches usually reddish). |
60+; corollas 5–7 mm, lobes yellow; anthers dark brown to black, appendages yellow. |
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Phyllaries | 24–30, lanceolate to lance-ovate, 8–13 × 2–5 mm, (margins sometimes ciliate) apices acute to short-acuminate or acuminate, abaxial faces sparsely hispid to glabrate. |
30–38 (loose, spreading), lanceolate to lance-ovate, 8–16 × 1.5–3 mm, (margins ciliate) apices acute to acuminate, abaxial faces usually ± strigose, rarely glabrate, not gland-dotted. |
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Heads | 1–3(–5). |
1–6. |
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Cypselae | 4–5 mm, glabrate; pappi of 2 aristate scales 1.7–2.5 mm plus 1–3 deltate scales 0.5–1.5 mm. |
3–5 mm, glabrate; pappi of 2 aristate scales 2.2–4.5 mm plus 0–2 lanceolate scales 0.5–1 mm. |
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Paleae | 7–9 mm, 3-toothed (apices purplish). |
8–12 mm, 3-toothed. |
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2n | = 34. |
= 34 (subspecies unknown). |
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Helianthus heterophyllus |
Helianthus nuttallii |
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Phenology | Flowering late summer–fall. | |||||||||
Habitat | Wet sandy soils | |||||||||
Elevation | 0–50+ m (0–200+ ft) | |||||||||
Distribution |
AL; FL; GA; LA; MS; NC; SC; TX
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AZ; CA; CO; IA; ID; KS; MN; MO; MT; ND; NE; NM; NV; OK; OR; SD; UT; WA; WY; AB; BC; MB; ON; QC; SK
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Discussion | Helianthus heterophyllus is found on the Atlantic and Gulf coastal plains. The cauline leaves are usually abruptly reduced relative to the basal leaves; individuals sometimes have relatively large cauline leaves. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 3 (3 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 21, p. 155. | FNA vol. 21, p. 166. | ||||||||
Parent taxa | Asteraceae > tribe Heliantheae > subtribe Helianthinae > Helianthus | Asteraceae > tribe Heliantheae > subtribe Helianthinae > Helianthus | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | H. elongatus | |||||||||
Name authority | Nuttall: J. Acad. Nat. Sci. Philadelphia 7: 74. (1834) | Torrey & A. Gray: Fl. N. Amer. 2: 324. (1842) | ||||||||
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