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cucumberleaf sunflower, weak sunflower

pineland sunflower, rayless sunflower, stiff sunflower

Habit Annuals or perennials, 30–200 cm (taprooted). Perennials, 50–100 cm (with crown buds).
Stems

decumbent to erect, glabrous, hirsute, or puberulent.

erect, distally densely hispid.

Leaves

mostly cauline; mostly alternate;

petioles 1–7 cm;

blades deltate-ovate, lance-ovate, or ovate, 2.5–14 × 1.8–13 cm, bases cordate to truncate or broadly cuneate, margins subentire to serrate, abaxial faces glabrate to hispid, not gland-dotted.

mostly basal; opposite;

petioles obscure (intergrading with blades);

blades obovate to orbiculate, 4.6–14.5 × 2.1–12 cm, bases broadly cuneate to rounded, margins entire or serrulate, abaxial faces strigoso-hispid, not gland-dotted (cauline usually much smaller, alternate distally).

Peduncles

9–50 cm.

10–20 cm.

Involucres

hemispheric, 10–22 mm diam.

shallowly hemispheric, (15–25 ×) 5–8 mm.

Ray florets

11–20;

laminae 12–23 mm.

0 or 2–8;

laminae (sometimes purplish) 1–2(–10) mm.

Disc florets

30+;

corollas 4.5–5 mm, lobes usually reddish, sometimes yellow;

anthers dark, appendages dark (style branches usually reddish, rarely yellow).

100–150+;

corollas 7–8 mm, lobes reddish;

anthers dark, appendages dark.

Phyllaries

20–30, lanceolate, 8–17 × 1–3 mm, apices acute to long-attenuate, abaxial faces glabrous or ± hispid, not gland-dotted.

(often dark purple) 25–33, lanceolate to ovate, 10–14 × 3–5 mm, apices acute to acuminate, abaxial faces hispid or glabrous.

Heads

1–3.

usually borne singly.

Cypselae

2.5–3.2 mm, glabrous or sparsely hairy;

pappi of 2 lanceolate or lance-linear scales 1.2–2.5 mm.

3–4 mm, glabrate;

pappi of 2 (often unequal) aristate scales 0.5–2.9 mm.

Paleae

7.5–8 mm, apices 3-toothed (middle teeth acuminate, usually glabrous or hispid, sometimes ± villous or bearded).

9–10 mm, subentire to 3-toothed (apices purplish, mucronate).

2n

= 34.

Helianthus debilis

Helianthus radula

Phenology Flowering fall.
Habitat Sandy, open pine barrens, flatwoods
Elevation 0–50+ m (0–200+ ft)
Distribution
from FNA
AL; CT; FL; GA; LA; MA; MD; MI; MS; NC; NH; NY; PA; RI; SC; TX; VA; VT; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; FL; GA; LA; MS; SC
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 5 (5 in the flora).

C. B. Heiser (1956) placed 8 subspecies in Helianthus debilis; he noted that alternative taxonomic treatments might recognize these in as many as three species, or expand the single species to include H. petiolaris. Later, Heiser et al. (1969) separated three of the subspecies as H. praecox. Isozyme data (R. P. Wain 1982, 1983; L. H. Rieseberg and M. F. Doyle 1989) show that all are closely related. Documented hybridization with H. annuus further complicates the situation. The treatment by Heiser et al. is followed here.

Helianthus debilis is adventive beyond the Atlantic and Gulf coasts of the United States.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems decumbent; peduncles 9–20(–22) cm
→ 2
1. Stems erect; peduncles (15–)20–50 cm
→ 3
2. Stems glabrous or puberulent; leaf blades serrulate or shallowly, regularly serrate, abaxial faces sparsely, if at all, gland-dotted
subsp. debilis
2. Stems hirsute; leaf blades deeply, irregularly serrate, abaxial faces densely gland-dotted
subsp. vestitus
3. Leaves 8–14 cm; peduncles (relatively slender) 20–40 cm; discs 10–15(–17) mm diam
subsp. silvestris
3. Leaves 2.5–9 cm; peduncles (not notably slender) 10–50 cm; discs 14–20 mm diam
→ 4
4. Leaf blades usually deeply, irregularly serrate; peduncles 10–25(–30) cm; ray lami-nae 12–20(–22) mm
subsp. tardiflorus
4. Leaf blades usually shallowly, regularly serrate; peduncles 25–50 cm; ray laminae (15–)20–23 mm
subsp. cucumerifolius
Source FNA vol. 21, p. 150. FNA vol. 21, p. 155.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Helianthinae > Helianthus Asteraceae > tribe Heliantheae > subtribe Helianthinae > Helianthus
Sibling taxa
H. agrestis, H. angustifolius, H. annuus, H. anomalus, H. argophyllus, H. arizonensis, H. atrorubens, H. bolanderi, H. californicus, H. carnosus, H. ciliaris, H. cusickii, H. decapetalus, H. deserticola, H. divaricatus, H. eggertii, H. exilis, H. floridanus, H. giganteus, H. glaucophyllus, H. gracilentus, H. grosseserratus, H. heterophyllus, H. hirsutus, H. laciniatus, H. laevigatus, H. longifolius, H. maximiliani, H. microcephalus, H. mollis, H. neglectus, H. niveus, H. nuttallii, H. occidentalis, H. paradoxus, H. pauciflorus, H. petiolaris, H. porteri, H. praecox, H. pumilus, H. radula, H. resinosus, H. salicifolius, H. schweinitzii, H. silphioides, H. simulans, H. smithii, H. strumosus, H. tuberosus, H. verticillatus, H. ×laetiflorus
H. agrestis, H. angustifolius, H. annuus, H. anomalus, H. argophyllus, H. arizonensis, H. atrorubens, H. bolanderi, H. californicus, H. carnosus, H. ciliaris, H. cusickii, H. debilis, H. decapetalus, H. deserticola, H. divaricatus, H. eggertii, H. exilis, H. floridanus, H. giganteus, H. glaucophyllus, H. gracilentus, H. grosseserratus, H. heterophyllus, H. hirsutus, H. laciniatus, H. laevigatus, H. longifolius, H. maximiliani, H. microcephalus, H. mollis, H. neglectus, H. niveus, H. nuttallii, H. occidentalis, H. paradoxus, H. pauciflorus, H. petiolaris, H. porteri, H. praecox, H. pumilus, H. resinosus, H. salicifolius, H. schweinitzii, H. silphioides, H. simulans, H. smithii, H. strumosus, H. tuberosus, H. verticillatus, H. ×laetiflorus
Subordinate taxa
H. debilis subsp. cucumerifolius, H. debilis subsp. debilis, H. debilis subsp. silvestris, H. debilis subsp. tardiflorus, H. debilis subsp. vestitus
Synonyms Rudbeckia radula
Name authority Nuttall: Trans. Amer. Philos. Soc., n. s. 7: 367. (1841) (Pursh) Torrey & A. Gray: Fl. N. Amer. 2: 321. (1842)
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