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cucumberleaf sunflower, weak sunflower

prairie sunflower, unknown

Habit Annuals or perennials, 30–200 cm (taprooted). Annuals, 40–200 cm.
Stems

decumbent to erect, glabrous, hirsute, or puberulent.

erect, usually densely canescent, hispid, or strigillose, rarely ± hirsute or glabrate.

Leaves

mostly cauline; mostly alternate;

petioles 1–7 cm;

blades deltate-ovate, lance-ovate, or ovate, 2.5–14 × 1.8–13 cm, bases cordate to truncate or broadly cuneate, margins subentire to serrate, abaxial faces glabrate to hispid, not gland-dotted.

mostly cauline; mostly alternate;

petioles 2–4 cm;

blades (often bluish green) lanceolate to deltate-ovate or ovate, 4–15 × 1–8 cm, bases subcordate or truncate to cuneate, margins entire or ± serrate, abaxial faces strigose, sparsely to densely, or not at all, gland-dotted.

Peduncles

9–50 cm.

4–15(–40) cm.

Involucres

hemispheric, 10–22 mm diam.

± hemispheric, 10–24 mm diam.

Ray florets

11–20;

laminae 12–23 mm.

10–30;

laminae 15–20 mm.

Disc florets

30+;

corollas 4.5–5 mm, lobes usually reddish, sometimes yellow;

anthers dark, appendages dark (style branches usually reddish, rarely yellow).

50–100+;

corollas 4.5–6 mm, lobes usually reddish, rarely yellow;

anthers reddish to purplish, appendages purplish (style branches reddish).

Phyllaries

20–30, lanceolate, 8–17 × 1–3 mm, apices acute to long-attenuate, abaxial faces glabrous or ± hispid, not gland-dotted.

14–25, lance-linear to lanceolate to lance-ovate, 10–14 × 1–4(–5) mm, (margins sometimes ciliate) apices short-attenuate, abaxial faces usually hispidulous, rarely sparsely hirsute to glabrate.

Heads

1–3.

1–5.

Cypselae

2.5–3.2 mm, glabrous or sparsely hairy;

pappi of 2 lanceolate or lance-linear scales 1.2–2.5 mm.

3–4.5 mm, ± villous;

pappi of 2 aristate scales 1.5–3 mm plus 0–2 erose scales 0.3–0.5 mm.

Paleae

7.5–8 mm, apices 3-toothed (middle teeth acuminate, usually glabrous or hispid, sometimes ± villous or bearded).

4.5–7.5 mm, 3-toothed, middle teeth ± ciliate or bearded, hairs whitish, 0.5–0.7 mm.

2n

= 34.

Helianthus debilis

Helianthus petiolaris

Distribution
from FNA
AL; CT; FL; GA; LA; MA; MD; MI; MS; NC; NH; NY; PA; RI; SC; TX; VA; VT; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AR; AZ; CA; CO; CT; DC; DE; IA; ID; IL; IN; KS; LA; MA; ME; MI; MN; MO; MT; NC; ND; NE; NJ; NM; NV; NY; OH; OK; OR; PA; SC; SD; TN; TX; UT; VA; WA; WI; WV; WY; AB; MB; ON; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 5 (5 in the flora).

C. B. Heiser (1956) placed 8 subspecies in Helianthus debilis; he noted that alternative taxonomic treatments might recognize these in as many as three species, or expand the single species to include H. petiolaris. Later, Heiser et al. (1969) separated three of the subspecies as H. praecox. Isozyme data (R. P. Wain 1982, 1983; L. H. Rieseberg and M. F. Doyle 1989) show that all are closely related. Documented hybridization with H. annuus further complicates the situation. The treatment by Heiser et al. is followed here.

Helianthus debilis is adventive beyond the Atlantic and Gulf coasts of the United States.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

Helianthus petiolaris is adventive beyond western North America.

A third subspecies has yet to be named at that rank in Helianthus petiolaris; it has been called H. petiolaris var. canescens A. Gray. It differs in having stems, leaves, and phyllaries densely canescent and abaxial faces of leaves densely gland-dotted. It is additionally characterized by peduncles usually ebracteate, phyllaries 1–2 mm wide, disc corolla throats gradually narrowed distal to slight, not densely hairy basal bulges, and 2n = 34. It flowers late spring through late summer and grows on sandy soils in open areas at (10–)1000–2300 m in Arizona, California, Nevada, New Mexico, and Texas and in Mexico. It was treated as H. niveus (Bentham) Brandegee subsp. canescens (A. Gray) Heiser by C. B. Heiser et al. (1969); molecular and morphologic data appear to favor a placement within H. petiolaris.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems decumbent; peduncles 9–20(–22) cm
→ 2
1. Stems erect; peduncles (15–)20–50 cm
→ 3
2. Stems glabrous or puberulent; leaf blades serrulate or shallowly, regularly serrate, abaxial faces sparsely, if at all, gland-dotted
subsp. debilis
2. Stems hirsute; leaf blades deeply, irregularly serrate, abaxial faces densely gland-dotted
subsp. vestitus
3. Leaves 8–14 cm; peduncles (relatively slender) 20–40 cm; discs 10–15(–17) mm diam
subsp. silvestris
3. Leaves 2.5–9 cm; peduncles (not notably slender) 10–50 cm; discs 14–20 mm diam
→ 4
4. Leaf blades usually deeply, irregularly serrate; peduncles 10–25(–30) cm; ray lami-nae 12–20(–22) mm
subsp. tardiflorus
4. Leaf blades usually shallowly, regularly serrate; peduncles 25–50 cm; ray laminae (15–)20–23 mm
subsp. cucumerifolius
1. Stems usually hispidulous to strigillose; peduncles usually bractless; phyllaries 3–5 mm wide; disc corollas: throats abruptly narrowed distal to densely hairy basal bulbs
subsp. petiolaris
1. Stems usually ± hispid; peduncles usually each with leafy bract subtending head; phyllaries 2–3.5 mm wide; disc corollas: throats gradually narrowed distal to slight, not densely hairy, basal bulges
subsp. fallax
Source FNA vol. 21, p. 150. FNA vol. 21, p. 152.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Helianthinae > Helianthus Asteraceae > tribe Heliantheae > subtribe Helianthinae > Helianthus
Sibling taxa
H. agrestis, H. angustifolius, H. annuus, H. anomalus, H. argophyllus, H. arizonensis, H. atrorubens, H. bolanderi, H. californicus, H. carnosus, H. ciliaris, H. cusickii, H. decapetalus, H. deserticola, H. divaricatus, H. eggertii, H. exilis, H. floridanus, H. giganteus, H. glaucophyllus, H. gracilentus, H. grosseserratus, H. heterophyllus, H. hirsutus, H. laciniatus, H. laevigatus, H. longifolius, H. maximiliani, H. microcephalus, H. mollis, H. neglectus, H. niveus, H. nuttallii, H. occidentalis, H. paradoxus, H. pauciflorus, H. petiolaris, H. porteri, H. praecox, H. pumilus, H. radula, H. resinosus, H. salicifolius, H. schweinitzii, H. silphioides, H. simulans, H. smithii, H. strumosus, H. tuberosus, H. verticillatus, H. ×laetiflorus
H. agrestis, H. angustifolius, H. annuus, H. anomalus, H. argophyllus, H. arizonensis, H. atrorubens, H. bolanderi, H. californicus, H. carnosus, H. ciliaris, H. cusickii, H. debilis, H. decapetalus, H. deserticola, H. divaricatus, H. eggertii, H. exilis, H. floridanus, H. giganteus, H. glaucophyllus, H. gracilentus, H. grosseserratus, H. heterophyllus, H. hirsutus, H. laciniatus, H. laevigatus, H. longifolius, H. maximiliani, H. microcephalus, H. mollis, H. neglectus, H. niveus, H. nuttallii, H. occidentalis, H. paradoxus, H. pauciflorus, H. porteri, H. praecox, H. pumilus, H. radula, H. resinosus, H. salicifolius, H. schweinitzii, H. silphioides, H. simulans, H. smithii, H. strumosus, H. tuberosus, H. verticillatus, H. ×laetiflorus
Subordinate taxa
H. debilis subsp. cucumerifolius, H. debilis subsp. debilis, H. debilis subsp. silvestris, H. debilis subsp. tardiflorus, H. debilis subsp. vestitus
H. petiolaris subsp. fallax, H. petiolaris subsp. petiolaris
Name authority Nuttall: Trans. Amer. Philos. Soc., n. s. 7: 367. (1841) Nuttall: J. Acad. Nat. Sci. Philadelphia 2: 115. (1821)
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