Helianthus bolanderi |
Helianthus eggertii |
|
---|---|---|
Bolander's sunflower, serpentine sunflower |
Eggert's sunflower |
|
Habit | Annuals, 60–150 cm. | Perennials, 30–200+ dm (rhizomatous). |
Stems | erect, hispid to hirsute. |
erect, glabrous (distinctively bluish, glacous). |
Leaves | mostly cauline; mostly alternate; petioles 1–4 cm; blades lance-linear or lance-ovate to ovate, 3–15 × 2–6 cm, bases cuneate to truncate, margins usually serrate, abaxial faces sparsely hirsute, gland-dotted. |
cauline; all or mostly opposite; sessile; blades (1-nerved) lanceolate to lance-ovate, 7–16.5 × 1.5–3.5 cm, bases cuneate, margins entire or serrulate, faces glabrous (abaxial) or strumose (adaxial), gland-dotted. |
Peduncles | 3–13 cm. |
1–4 cm. |
Involucres | hemispheric, 17–25 mm diam. |
hemispheric, 10–25 mm diam. |
Ray florets | 12–17; laminae 14–20 mm. |
10–18; laminae 15–20 mm. |
Disc florets | 75+ (discs usually 2+ cm diam.); corollas 5–7 mm, lobes usually reddish; anthers dark, appendages dark (style branches reddish). |
70+; corollas 5–7 mm, lobes yellow; anthers dark, appendages dark or ± yellowish. |
Phyllaries | 10–18, usually lanceolate to lance-ovate, 9–27 × (3–)3.5–5 mm (often surpassing discs), apices gradually attenuate, abaxial faces hirsute. |
30–38, lanceolate, 12–16 × 3.5–4.5 mm, (margins ciliate) apices acuminate, abaxial faces glabrate, not gland-dotted. |
Heads | 1–3. |
1–5. |
Cypselae | 3.5–4.5 mm, glabrate; pappi of 2 lanceolate scales 1.7–3 mm. |
4–6 mm, glabrous; pappi of 2 aristate scales 3–4 mm plus 0–1 deltate scales 0.5–1 mm. |
Paleae | 9.5–10.5 mm, 3-toothed (middle teeth subulate, surpassing discs, apices glabrous). |
9–11 mm, entire or ± 3-toothed (apices deltate). |
2n | = 34. |
= 102. |
Helianthus bolanderi |
Helianthus eggertii |
|
Phenology | Flowering summer–early fall. | Flowering late summer–fall. |
Habitat | Grassy, often disturbed sites | Open barrens, open oak-hickory woodlands |
Elevation | 10–1200 m (0–3900 ft) | 100–300 m (300–1000 ft) |
Distribution |
CA; OR
|
AL; KY; TN |
Discussion | Helianthus bolanderi and H. exilis form a closely related pair of sister species that share the distinctive feature of having the middle teeth of the paleae glabrous and relatively elongated, surpassing the disc florets. As treated here, H. bolanderi corresponds to the “valley weed race” (C. B. Heiser 1949; L. H. Rieseberg et al. 1988); it is separated from the “serpentine foothill race,” here recognized as H. exilis. Heiser proposed that H. bolanderi originated through hybridization between H. annuus and H. exilis; molecular studies by Rieseberg et al. do not support this scheme. In an ironic twist, it appears that H. bolanderi may be undergoing “genetic assimilation” through hybridization with H. annuus (S. E. Carney et al. 2000). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Helianthus eggertii is similar to H. strumosus and H. laevigatus; it is distinguished by the distinctive blue coloration of stems and leaves and by the leaves usually 1-nerved, in contrast to the 3-nerved condition typical for the genus. It is listed federally as an endangered species; R. L. Jones (1994) found it to be more common than was previously thought, and it is now in the process of being removed from the federal list. of conservation concern (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 21, p. 149. | FNA vol. 21, p. 159. |
Parent taxa | Asteraceae > tribe Heliantheae > subtribe Helianthinae > Helianthus | Asteraceae > tribe Heliantheae > subtribe Helianthinae > Helianthus |
Sibling taxa | ||
Name authority | A. Gray: Proc. Amer. Acad. Arts 6: 544. (1865) | Small: Fl. S.E. U.S., 1267, 1340. (1903) |
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