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blue milkwort

Habit Herbs, sometimes somewhat suffrutescent, (0.5–)0.8–6(10) dm. Herbs, perennial, or subshrubs, multi-stemmed [single].
Stems

usually erect, sometimes laxly so (rarely procumbent), hairs incurved (rarely glabrate).

prostrate or decumbent to erect, usually densely pubescent, rarely glabrate.

Leaves

blade ovate, elliptic, spatulate, or linear proximally, to elliptic, lanceolate, or linear distally (often longer than proximal ones), 4–45(–65) × 1.5–4(–6) mm, base usually acute, sometimes cuneate or obtuse, apex acute to acuminate or obtuse, surfaces not translucent-punctate, hairs incurved.

alternate; mostly sessile or subsessile (petiole to 2 mm in H. ovatifolia);

sometimes dimorphic (proximal different from distal);

blade surfaces not punctate (except translucent-punctate in H. macradenia and H. punctata), pubescent.

Inflorescences

terminal or leaf-opposed, racemes, loose, 2–12 × 0.9–1.4 cm;

peduncle 0.5–1.5 cm;

bracts linear to narrowly ovate.

terminal, axillary, or leaf-opposed, usually racemes, sometimes 1 or 2(–4)-flowered;

peduncle present or absent;

bracts deciduous.

Pedicels

1–5 mm, pubescent.

present.

Flowers

purplish, wings rarely yellow, keel distally yellow or yellowish green, 3.5–6 mm;

sepals lanceolate, 2.2–3.5 mm;

wings ovate to suborbiculate, 3.5–5.5 × 2.4–4 mm, sparsely pubescent;

keel 4–5.7 mm, pubescent.

usually purple, pink, yellow, greenish yellow, rarely whitish, wings sometimes cream, yellowish, or greenish, keel sometimes yellow or yellowish green distally, chasmogamous, 3–8 mm;

sepals deciduous or upper 1 sometimes tardily so or subpersistent (in H. macradenia and H. punctata), distinct, pubescent;

wings deciduous, 3.5–6(–7.5) mm, pubescent;

keel not beaked or crested, sometimes bluntly 3-lobed, pubescent or glabrate (in H. obscura);

stamens usually 8, sometimes 7 (in H. macradenia), not grouped;

ovary 2-loculed.

Fruits

capsules, dehiscent, margins not winged (except sometimes narrowly winged in H. barbeyana), not punctate (except translucent-punctate in H. macradenia and H. punctata), pubescent to glabrate or glabrous and ciliolate.

Capsules

oblong, oblong-ovoid, broadly ellipsoid, or subglobose, 5–10(–11) ×4–7 mm, not translucent-punctate, glabrous and ciliolate, margins sometimes narrowly winged.

Seeds

4–4.5 mm;

aril 0.6–1 mm, lobes vestigial or to 1/4 length of seed.

pubescent, arillate.

2n

= 30.

Hebecarpa barbeyana

Hebecarpa

Phenology Flowering spring–late fall.
Habitat Gravelly or rocky soils, mostly limestone, or igneous or gypsum, arid or semi-arid scrub, open woodlands, chaparral.
Elevation 400–2400(–2800) m. (1300–7900(–9200) ft.)
Distribution
from FNA
AZ; NM; TX; Mexico (Coahuila, Durango, Guanajuato, Jalisco, Nuevo León, Querétaro, San Luis Potosí, Veracruz, Zacatecas)
[BONAP county map]
sw United States; sc United States; Mexico; Central America; South America; West Indies
[BONAP county map]
Discussion

Hebecarpa barbeyana is very similar to 7. H. rectipilis; see that species for comments.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 40–70 (7 in the flora).

Most species of Hebecarpa are Mexican, the center of diversity for this group. The seven in the flora area occur mostly in arid and semi-arid areas of the southwestern United States, representing contiguous northern expansions of their ranges in Mexico. A few species range into Central America and Andean South America, but they are not the basalmost species, suggesting that they have arrived in these regions secondarily.

Several species complexes in this group are in serious need of revision, with overly fine, historical splitting (often based on few specimens); thus the total number of species is probably fewer than 50.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blade surfaces and capsules densely translucent-punctate from internal cavities appearing as pustular glands 0.2–0.4 mm diam.
→ 2
2. Leaf blades linear-oblong to oblong-lanceolate, narrowly ovate, or linear.
H. macradenia
2. Leaf blades usually obovate, rarely suborbiculate to broadly elliptic.
H. punctata
1. Leaf blade surfaces and capsules not translucent-punctate.
→ 3
3. Capsules densely pubescent to glabrate.
H. obscura
3. Capsules glabrous, margins ciliolate.
→ 4
4. Stems and leaves with incurved hairs.
→ 5
5. Middle and distal leaf blades usually 1.5–4(–6) mm wide, elliptic, lanceolate, or linear; capsules 5–10(–11) mm.
H. barbeyana
5. Middle and distal leaf blades 4–7 mm wide, ovate, elliptic, or obovate; capsules 9–12 mm.
H. palmeri
4. Stems and leaves with spreading hairs.
→ 6
6. Leaf blades ovate to ovate-oblong or elliptic; capsules 8.5–13 mm.
H. ovatifolia
6. Leaf blades ovate, elliptic, spatulate, or linear proximally, elliptic, lanceolate, or linear distally (often longer than proximal ones); capsules 6–8 mm.
H. rectipilis
Source FNA vol. 10. FNA vol. 10.
Parent taxa Polygalaceae > Hebecarpa Polygalaceae
Sibling taxa
H. macradenia, H. obscura, H. ovatifolia, H. palmeri, H. punctata, H. rectipilis
Subordinate taxa
H. barbeyana, H. macradenia, H. obscura, H. ovatifolia, H. palmeri, H. punctata, H. rectipilis
Synonyms Polygalabarbeyana chodat, P. longa, P. racemosa, P. reducta Polygala section hebecarpa
Name authority (Chodat) J. R. Abbott: J. Bot. Res. Inst. Texas 5: 134. (2011) (Chodat) J. R. Abbott: J. Bot. Res. Inst. Texas 5: 134. (2011)
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