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subalpine gumweed

Howell's gumweed

Habit Biennials or perennials (perhaps flowering first year), 15–30(–60) cm. Perennials, (25–)60–90+ cm.
Stems

erect, usually reddish, sometimes stramineous, glabrous.

erect, stramineous to red-brown, stipitate-glandular (at least distally).

Cauline leaf

blades spatulate or oblanceolate to oblong or lanceolate, 15–70 mm, lengths 3–5(–7) times widths, bases ± cuneate or clasping, margins usually dentate or serrate (teeth apiculate), sometimes entire, apices rounded or obtuse to acute, faces glabrous (or, near margins, scabridulous) and sparsely to densely gland-dotted.

blades spatulate or oblong to oblanceolate or lanceolate, 25–60(–90) mm, lengths 3–5(–7) times widths, bases cuneate to ± clasping, margins entire or serrate to denticulate (teeth apiculate), apices obtuse to acute, faces usually finely stipitate-glandular, sometimes glabrous (or scabridulous near margins) and gland-dotted.

Involucres

broadly urceolate to globose, 8–11 × 10–20 mm.

broadly urceolate to globose, 8–15 × 12–20(–30) mm.

Ray florets

18–27;

laminae 10–15 mm.

20–30+;

laminae 8–12 mm.

Phyllaries

in 5–6 series, linear or lanceolate to oblong-lanceolate, apices looped to hooked (outer) or recurved to nearly straight, terete to subulate or acuminate, slightly to moderately resinous.

in 6–9 series, spreading to appressed, linear or linear-attenuate to lance-linear, apices mostly looped to hooked (inner sometimes recurved to straight), ± terete, moderately to strongly resinous.

Heads

usually in open, corymbiform to paniculiform arrays, seldom borne singly.

in open to crowded, corymbiform to paniculiform arrays.

Cypselae

whitish or stramineous, 2.5–5 mm, apices smooth or weakly coronate, faces smooth or finely furrowed;

pappi of 4–8 straight, barbellate to barbellulate bristles or setiform awns 2.5–5.5 mm, shorter than disc corollas.

whitish to stramineous, 4–5.5 mm, apices ± coronate, faces striate or furrowed;

pappi of 2 straight to curled, usually smooth, sometimes barbellulate, subulate scales 2.5–4+ mm, shorter than disc corollas.

2n

= 12.

Grindelia subalpina

Grindelia howellii

Phenology Flowering Jul–Aug. Flowering Jul–Sep.
Habitat Dry, open, sandy or gravelly slopes, ridges, valleys, stream banks, waste grounds Disturbed sites, grasslands, forest openings
Elevation 1600–2900 m (5200–9500 ft) 900–1300 m (3000–4300 ft)
Distribution
from FNA
CO; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
ID; MT
[WildflowerSearch map]
Discussion

Of conservation concern.

But for the usually stipitate-glandular indument of the stems and leaves, plants called Grindelia howellii are very much like plants that have been called G. nana forma brownii Steyermark, G. nana forma longisquama Steyermark, and G. paysonorum H. St. John [= G. nana var. paysonorum (H. St. John) Steyermark], all typified by specimens included here in G. hirsutula. In some plants of G. howellii, the glands on the leaves range from stipitate to ± embedded (e.g., Pierce 1146, ID). Taxonomic rank for plants that have been called G. howellii should be reconsidered.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 20, p. 434. FNA vol. 20, p. 431.
Parent taxa Asteraceae > tribe Astereae > Grindelia Asteraceae > tribe Astereae > Grindelia
Sibling taxa
G. adenodonta, G. arizonica, G. ciliata, G. decumbens, G. fraxinipratensis, G. grandiflora, G. havardii, G. hirsutula, G. howellii, G. integrifolia, G. lanceolata, G. microcephala, G. oölepis, G. oxylepis, G. pusilla, G. scabra, G. squarrosa
G. adenodonta, G. arizonica, G. ciliata, G. decumbens, G. fraxinipratensis, G. grandiflora, G. havardii, G. hirsutula, G. integrifolia, G. lanceolata, G. microcephala, G. oölepis, G. oxylepis, G. pusilla, G. scabra, G. squarrosa, G. subalpina
Synonyms G. erecta, G. subalpina var. erecta
Name authority Greene: Pittonia 3: 297. (1898) Steyermark: Ann. Missouri Bot. Gard. 21: 549, fig. 30. (1934)
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