Grimmia trichophylla |
Grimmia ramondii |
|
---|---|---|
grimmia dry rock moss, hair-point grimmia |
spreading-leaf grimmia |
|
Habit | Plants in dense to loose patches, yellowish green to dark green. | Plants in robust, loose, arched-ascending, readily disintegrating patches, brownish green distally, blackish proximally. |
Stems | 2–4 cm, central strand present. |
5–10 cm, central strand absent. |
Gemmae | clusters occasionally present in distal leaf axils. |
absent. |
Leaves | loosely appressed, slightly twisted when dry, erecto-patent when moist, lanceolate to oblong-lanceolate, tapering to acute apex, 2–3.5 × 0.3–0.4 mm, usually sharply keeled, margins recurved on one or both sides, plane to erect distally, awns variable, short to long, smooth to denticulate, not conspicuously flattened at base, costa firm, projecting on abaxial side; basal juxtacostal laminal cells long-rectangular (rarely short-rectangular), ± nodulose, thick-walled; basal marginal laminal cells short- to long-rectangular, with thickened transverse walls; medial laminal cells quadrate to short-rectangular, slightly sinuose, thick-walled; distal laminal cells 1-stratose, occasionally with 2-stratose ridges. |
loosely appressed when dry, patent when moist, broadly ovate-lanceolate, tapering to an acute, slightly toothed apex, 2–3 × 0.4–0.6 mm, keeled, margins recurved on both sides, awns absent, costa stout, with two abaxial lamellae (or wings) forming parallel ridges along its length; basal juxtacostal laminal cells yellow, linear, sinuose, thick-walled; basal marginal laminal cells quadrate to short-rectangular, slightly thick-walled; medial laminal cells quadrate to rectangular, sinuous, thick-walled; distal laminal cells 1-stratose, margins 2-stratose. |
Seta | arcuate, 2–4 mm. |
cygneous to arcuate at maturity, flexuose when old, 3–5 mm. |
Sexual condition | dioicous. |
dioicous. |
Capsule | occasionally present, exserted, oblong-ovoid, yellowish green to stramineous, striate when dry, exothecial cells thin-walled, annulus present, operculum rostrate, peristome teeth yellowish, papillose, deeply split and perforated. |
occasionally present, exserted, obloid, yellowish green to yellowish brown, plicate when empty, exothecial cells thin-walled, annulus present, operculum rostrate, peristome teeth purple, divided nearly to base into two partly adhering segments, papillose. |
Calyptra | mitrate. |
mitrate. |
Grimmia trichophylla |
Grimmia ramondii |
|
Habitat | Dry, acidic rock | Dry to damp acidic rock |
Elevation | moderate to high elevations (200-2000 m) [moderate to high elevations (700-6600 ft)] | low to high elevations (0-2000 m) [low to high elevations (0-6600 ft)] |
Distribution |
AZ; CA; CO; ID; ME; MO; MT; NV; OK; OR; SD; UT; VT; WA; WY; HI; BC; Mexico; Eurasia; Australia
|
AK; CA; ID; MI; MT; OR; WA; AB; BC; NF; NT; Greenland; Europe; Asia (China, Japan); n Africa
|
Discussion | In North America, Grimmia trichophylla is principally a lowland species, occurring in the mountains up to about 1000 m., rarely higher. In the Southern Hemisphere, it may be found up to 4000 m. In New Zealand, the species is common, and in contrast to G. trichophylla in North America, frequently bears capsules. The New Zealand plants are usually smaller than American specimens, and the leaves are frequently contorted. The nearly cosmopolitan G. trichophylla has many phenotypes, and numerous subspecies and varieties have been described. In damp and shaded habitats, the awns may be short, just as in dry unfavorable habitats at high altitudes, where stunted specimens may occur with small, short leaves and reduced awns, or even with muticous leaves. Grimmia trichophylla has frequently been confused with related species such as 36. G. muehlenbeckii and 34. G. lisae (see discussions thereunder for identification details). Robust forms of G. trichophylla have been mistaken for G. austrofunalis (H. C. Greven 1997, 2003), which does not occur in North America. Although some of those plants have leaves of equal length along the stem, characteristic of G. austrofunalis, they also have both leaf margins recurved, and the medial and outer basal laminal cells are longer and more robust than in that species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Grimmia ramondii occurs near sea level in arctic areas and along the West Coast. In the latter region, however, it is more common in mid-elevation montane sites and may also be found above the tree line. It differs from most Grimmia species by its lack of awns, winged costa, and tall, loose growth form. At first sight, G. ramondii has the habit of a Racomitrium. However, the sporophyte has an arcuate rather than straight seta, and the capsules are striate rather than smooth. Because of its habit and the absence of a central strand, K. G. Limpricht ([1885–]1890–1903, vol. 1) placed it, together with G. hartmanii and G. atrata, in Dryptodon, a genus intermediate between Grimmia and Racomitrium. There are many other Grimmia species, however, without a central strand, and G. hartmanii and G. atrata have already been removed from Dryptodon, so there is no reason to maintain that monotypic genus. Grimmia ramondii may be confused with G. hartmanii and G. elatior. However, the lack of awns, the Racomitrioideae-like areolation, and the distinct costal wings distinguish it easily from those species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 257. | FNA vol. 27, p. 256. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Pterigynandrum ramondii, Dryptodon patens, G. curvata, Racomitrium patens | |
Name authority | Greville: Fl. Edin., 235. (1824) | (Lamarck & De Candolle) Margadant: Lindbergia 1: 128. (1972) |
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