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grimmia dry rock moss, hair-point grimmia

grimmia dry rock moss

Habit Plants in dense to loose patches, yellowish green to dark green. Plants in dense cushions to hoary tufts, dark green to brown.
Stems

2–4 cm, central strand present.

0.3–0.5(–1) cm.

Gemmae

clusters occasionally present in distal leaf axils.

Leaves

loosely appressed, slightly twisted when dry, erecto-patent when moist, lanceolate to oblong-lanceolate, tapering to acute apex, 2–3.5 × 0.3–0.4 mm, usually sharply keeled, margins recurved on one or both sides, plane to erect distally, awns variable, short to long, smooth to denticulate, not conspicuously flattened at base, costa firm, projecting on abaxial side;

basal juxtacostal laminal cells long-rectangular (rarely short-rectangular), ± nodulose, thick-walled;

basal marginal laminal cells short- to long-rectangular, with thickened transverse walls;

medial laminal cells quadrate to short-rectangular, slightly sinuose, thick-walled;

distal laminal cells 1-stratose, occasionally with 2-stratose ridges.

oblong-ovate, 1–1.7 × 0.4–0.8 mm, concave-keeled, awn 0.3–1 mm;

basal juxtacostal laminal cells quadrate to short-rectangular, straight, thin-walled;

basal marginal laminal cells quadrate to short-rectangular, straight, thin-walled;

medial laminal cells quadrate to short-rectangular, slightly sinuose, slightly thick-walled;

distal laminal cells 1-stratose, marginal cells 1–2-stratose.

Seta

arcuate, 2–4 mm.

sigmoid, 0.2–0.3 mm.

Sexual condition

dioicous.

gonioautoicous.

Capsule

occasionally present, exserted, oblong-ovoid, yellowish green to stramineous, striate when dry, exothecial cells thin-walled, annulus present, operculum rostrate, peristome teeth yellowish, papillose, deeply split and perforated.

usually present, exothecial cells thin-walled, annulus absent, operculum mammillate, peristome present, fully developed, perforated and split in distal half.

Calyptra

mitrate.

Grimmia trichophylla

Grimmia plagiopodia

Habitat Dry, acidic rock Exposed calcareous sandstone, limestone, occasionally concrete, and glacio-lacustrine silt
Elevation moderate to high elevations (200-2000 m) (moderate to high elevations (700-6600 ft)) low to high elevations (50-2400 m) (low to high elevations (200-7900 ft))
Distribution
from FNA
AZ; CA; CO; ID; ME; MO; MT; NV; OK; OR; SD; UT; VT; WA; WY; HI; BC; Mexico; Eurasia; Australia
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from FNA
AK; CA; CO; IA; ID; IL; MN; MT; ND; NE; NM; NV; SD; UT; WI; WY; AB; BC; NT; NU; ON; SK; South America; Pacific Islands (New Zealand); Greenland; Eurasia; Antarctic
Discussion

In North America, Grimmia trichophylla is principally a lowland species, occurring in the mountains up to about 1000 m., rarely higher. In the Southern Hemisphere, it may be found up to 4000 m. In New Zealand, the species is common, and in contrast to G. trichophylla in North America, frequently bears capsules. The New Zealand plants are usually smaller than American specimens, and the leaves are frequently contorted. The nearly cosmopolitan G. trichophylla has many phenotypes, and numerous subspecies and varieties have been described. In damp and shaded habitats, the awns may be short, just as in dry unfavorable habitats at high altitudes, where stunted specimens may occur with small, short leaves and reduced awns, or even with muticous leaves. Grimmia trichophylla has frequently been confused with related species such as 36. G. muehlenbeckii and 34. G. lisae (see discussions thereunder for identification details). Robust forms of G. trichophylla have been mistaken for G. austrofunalis (H. C. Greven 1997, 2003), which does not occur in North America. Although some of those plants have leaves of equal length along the stem, characteristic of G. austrofunalis, they also have both leaf margins recurved, and the medial and outer basal laminal cells are longer and more robust than in that species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Grimmia plagiopodia has a widespread and continuous distribution on calcareous rock across the northern Great Plains, reaching as far east as Illinois. It is rare in eastern North America, with a disjunct site in southern Ontario. In the west it reaches into the mountains on limestone and basic sandstone deposits, but its continuous range does not extend west of a line from Utah to south-central British Columbia. There is a disjunct location near Carson City, Nevada and Lake Tahoe, California. In the Arctic it is known from a few scattered localities extending from northwestern Greenland and nearby Ellesmere Island to the North Slope of Alaska. Compared to G. anodon, G. plagiopodia tends to occupy more prairie-like sites and is typically found at lower elevations. Commonly fertile, it is recognized by its immersed, peristomate capsule on a sigmoid seta with fully-developed teeth that are perforated and split distally. Grimmia americana is similar but has a short, straight to arcuate seta and a large annulus. The other widespread species in the group, G. anodon, has an annulus and is gymnostomous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 257. FNA vol. 27, p. 231.
Parent taxa Grimmiaceae > subfam. Grimmioideae > Grimmia > subg. Rhabdogrimmia Grimmiaceae > subfam. Grimmioideae > Grimmia > subg. Grimmia
Sibling taxa
G. alpestris, G. americana, G. anodon, G. anomala, G. arizonae, G. atrata, G. attenuata, G. brittoniae, G. caespiticia, G. crinitoleucophaea, G. donniana, G. elatior, G. elongata, G. funalis, G. hamulosa, G. hartmanii, G. incurva, G. laevigata, G. leibergii, G. lesherae, G. lisae, G. longirostris, G. mariniana, G. mollis, G. montana, G. moxleyi, G. muehlenbeckii, G. nevadensis, G. olneyi, G. orbicularis, G. ovalis, G. pilifera, G. plagiopodia, G. pulvinata, G. ramondii, G. reflexidens, G. serrana, G. sessitana, G. shastae, G. teretinervis, G. torquata, G. unicolor
G. alpestris, G. americana, G. anodon, G. anomala, G. arizonae, G. atrata, G. attenuata, G. brittoniae, G. caespiticia, G. crinitoleucophaea, G. donniana, G. elatior, G. elongata, G. funalis, G. hamulosa, G. hartmanii, G. incurva, G. laevigata, G. leibergii, G. lesherae, G. lisae, G. longirostris, G. mariniana, G. mollis, G. montana, G. moxleyi, G. muehlenbeckii, G. nevadensis, G. olneyi, G. orbicularis, G. ovalis, G. pilifera, G. pulvinata, G. ramondii, G. reflexidens, G. serrana, G. sessitana, G. shastae, G. teretinervis, G. torquata, G. trichophylla, G. unicolor
Synonyms G. brandegeei
Name authority Greville: Fl. Edin., 235. (1824) Hedwig: Sp. Musc. Frond., 78, plate 15, figs. 6–13. (1801)
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