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grimmia dry rock moss, hair-point grimmia

Habit Plants in dense to loose patches, yellowish green to dark green. Plants in dense cushions, blackish green.
Stems

2–4 cm, central strand present.

0.5–0.7(–1) cm, central strand weak.

Gemmae

clusters occasionally present in distal leaf axils.

Leaves

loosely appressed, slightly twisted when dry, erecto-patent when moist, lanceolate to oblong-lanceolate, tapering to acute apex, 2–3.5 × 0.3–0.4 mm, usually sharply keeled, margins recurved on one or both sides, plane to erect distally, awns variable, short to long, smooth to denticulate, not conspicuously flattened at base, costa firm, projecting on abaxial side;

basal juxtacostal laminal cells long-rectangular (rarely short-rectangular), ± nodulose, thick-walled;

basal marginal laminal cells short- to long-rectangular, with thickened transverse walls;

medial laminal cells quadrate to short-rectangular, slightly sinuose, thick-walled;

distal laminal cells 1-stratose, occasionally with 2-stratose ridges.

ovate-lanceolate, 1.5–2 mm, both margins plane, incurved distally, intermarginal bands absent, awn 0.4–0.8 mm, costal transverse section prominent, semicircular;

basal juxtacostal laminal cells short-rectangular, straight, thin-walled;

basal marginal laminal cells short-rectangular, straight, with thick transverse and thin lateral walls, not hyaline;

medial laminal cells quadrate to rounded, straight, thin-walled;

distal laminal cells 2-stratose.

Seta

arcuate, 2–4 mm.

straight, 0.9–1.1 mm.

Sexual condition

dioicous.

Capsule

occasionally present, exserted, oblong-ovoid, yellowish green to stramineous, striate when dry, exothecial cells thin-walled, annulus present, operculum rostrate, peristome teeth yellowish, papillose, deeply split and perforated.

commonly present, emergent, chestnut brown, ovate with distinct constriction below rim, exothecial cells rectangular, thick-walled, stomata absent, annulus of 1 row of quadrate, thick-walled cells, operculum conic, peristome absent.

Calyptra

mitrate.

cucullate.

Perichaetial

leaves not enlarged.

Sexuality

dioicous.

Grimmia trichophylla

Grimmia nevadensis

Habitat Dry, acidic rock Acidic sedimentary rocks
Elevation moderate to high elevations (200-2000 m) [moderate to high elevations (700-6600 ft)] moderate to high elevations (1900-2500 m) [moderate to high elevations (6200-8200 ft)]
Distribution
from FNA
AZ; CA; CO; ID; ME; MO; MT; NV; OK; OR; SD; UT; VT; WA; WY; HI; BC; Mexico; Eurasia; Australia
[WildflowerSearch map]
from FNA
CA; NV
Discussion

In North America, Grimmia trichophylla is principally a lowland species, occurring in the mountains up to about 1000 m., rarely higher. In the Southern Hemisphere, it may be found up to 4000 m. In New Zealand, the species is common, and in contrast to G. trichophylla in North America, frequently bears capsules. The New Zealand plants are usually smaller than American specimens, and the leaves are frequently contorted. The nearly cosmopolitan G. trichophylla has many phenotypes, and numerous subspecies and varieties have been described. In damp and shaded habitats, the awns may be short, just as in dry unfavorable habitats at high altitudes, where stunted specimens may occur with small, short leaves and reduced awns, or even with muticous leaves. Grimmia trichophylla has frequently been confused with related species such as 36. G. muehlenbeckii and 34. G. lisae (see discussions thereunder for identification details). Robust forms of G. trichophylla have been mistaken for G. austrofunalis (H. C. Greven 1997, 2003), which does not occur in North America. Although some of those plants have leaves of equal length along the stem, characteristic of G. austrofunalis, they also have both leaf margins recurved, and the medial and outer basal laminal cells are longer and more robust than in that species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Grimmia nevadensis is a rare but locally abundant endemic, known only from the Sierra Nevada Mountains. Fertile specimens of it can be separated from G. mariniana by the absence of peristome teeth and by the chestnut brown capsules. Gametophytically, specimens of G. nevadensis contrast with those of G. mariniana by having a uniform basal areolation of short-rectangular cells with thin lateral walls, and margins not hyaline. If fertile, the immersed, eperistomate capsules will separate G. nevadensis from G. ovalis, G. alpestris, and G. montana. Sterile specimens can be separated from G. alpestris by the absence of bulging cells, and the uniformly short-rectangular, thin-walled basal laminal cells. While both G. nevadensis and G. montana have plane to incurved distal margins and both lack bulging laminal cells, the uniformly short-rectangular, thin-walled basal laminal cells and concave leaves will identify G. nevadensis. The latter is easily separated from G. arizonae and G. pilifera, in contrast with which it lacks stomata and peristome teeth, and its concave leaves with plane to incurved margins differ markedly from the keeled leaves with recurved leaf margins of the others.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 257. Treatment authors: Roxanne I. Hastings, Henk C. Greven. FNA vol. 27, p. 245. Treatment authors: Roxanne I. Hastings, Henk C. Greven.
Parent taxa Grimmiaceae > subfam. Grimmioideae > Grimmia > subg. Rhabdogrimmia Grimmiaceae > subfam. Grimmioideae > Grimmia > subg. Litoneuron
Sibling taxa
G. alpestris, G. americana, G. anodon, G. anomala, G. arizonae, G. atrata, G. attenuata, G. brittoniae, G. caespiticia, G. crinitoleucophaea, G. donniana, G. elatior, G. elongata, G. funalis, G. hamulosa, G. hartmanii, G. incurva, G. laevigata, G. leibergii, G. lesherae, G. lisae, G. longirostris, G. mariniana, G. mollis, G. montana, G. moxleyi, G. muehlenbeckii, G. nevadensis, G. olneyi, G. orbicularis, G. ovalis, G. pilifera, G. plagiopodia, G. pulvinata, G. ramondii, G. reflexidens, G. serrana, G. sessitana, G. shastae, G. teretinervis, G. torquata, G. unicolor
G. alpestris, G. americana, G. anodon, G. anomala, G. arizonae, G. atrata, G. attenuata, G. brittoniae, G. caespiticia, G. crinitoleucophaea, G. donniana, G. elatior, G. elongata, G. funalis, G. hamulosa, G. hartmanii, G. incurva, G. laevigata, G. leibergii, G. lesherae, G. lisae, G. longirostris, G. mariniana, G. mollis, G. montana, G. moxleyi, G. muehlenbeckii, G. olneyi, G. orbicularis, G. ovalis, G. pilifera, G. plagiopodia, G. pulvinata, G. ramondii, G. reflexidens, G. serrana, G. sessitana, G. shastae, G. teretinervis, G. torquata, G. trichophylla, G. unicolor
Name authority Greville: Fl. Edin., 235. (1824) Greven: Bryologist 105: 273, fig. 1. 2002 (as nevadense),
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