Grimmia trichophylla |
Grimmia lisae |
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grimmia dry rock moss, hair-point grimmia |
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Habit | Plants in dense to loose patches, yellowish green to dark green. | Plants in dense to loose tufts, olivaceous, brownish to blackish proximally. |
Stems | 2–4 cm, central strand present. |
1–4 cm, central strand present. |
Gemmae | clusters occasionally present in distal leaf axils. |
in clusters, multicellular, occasionally present in leaf axils. |
Leaves | loosely appressed, slightly twisted when dry, erecto-patent when moist, lanceolate to oblong-lanceolate, tapering to acute apex, 2–3.5 × 0.3–0.4 mm, usually sharply keeled, margins recurved on one or both sides, plane to erect distally, awns variable, short to long, smooth to denticulate, not conspicuously flattened at base, costa firm, projecting on abaxial side; basal juxtacostal laminal cells long-rectangular (rarely short-rectangular), ± nodulose, thick-walled; basal marginal laminal cells short- to long-rectangular, with thickened transverse walls; medial laminal cells quadrate to short-rectangular, slightly sinuose, thick-walled; distal laminal cells 1-stratose, occasionally with 2-stratose ridges. |
erect and appressed when dry, recurved to squarrose when moist, broadly lanceolate, tapering to an acute apex, 1.5–2.5 × 0.4–0.6 mm, keeled, margins recurved on one or both sides, awns absent to rather long, stout and denticulate, costa reniform, projecting on dorsal side, median layer of stereids present; basal juxtacostal laminal cells short-rectangular to occasionally elongate, straight to slightly sinuose, thin- to thick-walled; basal marginal laminal cells quadrate to short-rectangular, thickened transverse walls; medial laminal cells oblate to rounded-quadrate, straight, thick-walled; distal laminal cells 1-stratose with 2-stratose ridges, margins 2-stratose. |
Seta | arcuate, 2–4 mm. |
arcuate, 3–4.5 mm. |
Sexual condition | dioicous. |
dioicous. |
Capsule | occasionally present, exserted, oblong-ovoid, yellowish green to stramineous, striate when dry, exothecial cells thin-walled, annulus present, operculum rostrate, peristome teeth yellowish, papillose, deeply split and perforated. |
occasionally present, exserted, ovoid, brown, shiny, weakly striate, exothecial cells thin-walled, annulus present, operculum rostrate, peristome teeth orange, fully-developed to irregularly cleft at apex, papillose. |
Calyptra | mitrate. |
mitrate. |
Grimmia trichophylla |
Grimmia lisae |
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Habitat | Dry, acidic rock | Dry acidic to basic rock |
Elevation | moderate to high elevations (200-2000 m) [moderate to high elevations (700-6600 ft)] | low to moderate elevations (60-1000 m) [low to moderate elevations (200-3300 ft)] |
Distribution |
AZ; CA; CO; ID; ME; MO; MT; NV; OK; OR; SD; UT; VT; WA; WY; HI; BC; Mexico; Eurasia; Australia
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CA; OR; WA; HI; BC; Mexico; Eurasia; n Africa |
Discussion | In North America, Grimmia trichophylla is principally a lowland species, occurring in the mountains up to about 1000 m., rarely higher. In the Southern Hemisphere, it may be found up to 4000 m. In New Zealand, the species is common, and in contrast to G. trichophylla in North America, frequently bears capsules. The New Zealand plants are usually smaller than American specimens, and the leaves are frequently contorted. The nearly cosmopolitan G. trichophylla has many phenotypes, and numerous subspecies and varieties have been described. In damp and shaded habitats, the awns may be short, just as in dry unfavorable habitats at high altitudes, where stunted specimens may occur with small, short leaves and reduced awns, or even with muticous leaves. Grimmia trichophylla has frequently been confused with related species such as 36. G. muehlenbeckii and 34. G. lisae (see discussions thereunder for identification details). Robust forms of G. trichophylla have been mistaken for G. austrofunalis (H. C. Greven 1997, 2003), which does not occur in North America. Although some of those plants have leaves of equal length along the stem, characteristic of G. austrofunalis, they also have both leaf margins recurved, and the medial and outer basal laminal cells are longer and more robust than in that species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Grimmia lisae is a thermophilous species with a preference for subtropical coastal areas. In North America, it occurs along the west coast, from Vancouver Island south to Mexico. From that region, it has been described frequently as a new species. It is closely related to G. trichophylla, but is distinguished by somewhat shorter and broader leaves that are straight and appressed when dry and recurved to squarrose when moist, and by a reniform costa. Furthermore, it is characterized by a grass-green mid leaf areolation with small, rounded, frequently oblate cells with straight walls. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 257. | FNA vol. 27, p. 253. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | G. ancistrodes, G. arcuatifolia, G. californica, G. canadensis, G. flettii | |
Name authority | Greville: Fl. Edin., 235. (1824) | De Notaris: Muscol. Ital. Spic., 15. (1837) |
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