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grimmia dry rock moss, hair-point grimmia

American dry rock moss

Habit Plants in dense to loose patches, yellowish green to dark green. Plants in hoary tufts, dark green to brown.
Stems

2–4 cm, central strand present.

0.5–1.5 cm.

Gemmae

clusters occasionally present in distal leaf axils.

Leaves

loosely appressed, slightly twisted when dry, erecto-patent when moist, lanceolate to oblong-lanceolate, tapering to acute apex, 2–3.5 × 0.3–0.4 mm, usually sharply keeled, margins recurved on one or both sides, plane to erect distally, awns variable, short to long, smooth to denticulate, not conspicuously flattened at base, costa firm, projecting on abaxial side;

basal juxtacostal laminal cells long-rectangular (rarely short-rectangular), ± nodulose, thick-walled;

basal marginal laminal cells short- to long-rectangular, with thickened transverse walls;

medial laminal cells quadrate to short-rectangular, slightly sinuose, thick-walled;

distal laminal cells 1-stratose, occasionally with 2-stratose ridges.

oblong-ovate to oblong-lanceolate, 1.5–2 × 0.5–0.9 mm, concave, awn to 1 mm;

basal juxtacostal laminal cells quadrate to short-rectangular, straight, thick-walled;

basal marginal laminal cells quadrate to long-rectangular, straight, thick-walled, medial laminal cells rounded-quadrate, slightly thick-walled;

distal laminal cells 2-stratose, marginal cells 2-stratose.

Seta

arcuate, 2–4 mm.

straight to arcuate, 0.6–1 mm.

Sexual condition

dioicous.

gonioautoicous.

Capsule

occasionally present, exserted, oblong-ovoid, yellowish green to stramineous, striate when dry, exothecial cells thin-walled, annulus present, operculum rostrate, peristome teeth yellowish, papillose, deeply split and perforated.

usually present, exothecial cells thick-walled, annulus of 2–3 rows of rectangular, thin-walled cells, revoluble, operculum rostellate, peristome present, fully developed, perforated and split in most distal part, weakly papillose.

Calyptra

mitrate.

Grimmia trichophylla

Grimmia americana

Habitat Dry, acidic rock Calcareous rock
Elevation moderate to high elevations (200-2000 m) [moderate to high elevations (700-6600 ft)] moderate to high elevations (ca. 1600 m) [moderate to high elevations (ca. 5200 ft)]
Distribution
from FNA
AZ; CA; CO; ID; ME; MO; MT; NV; OK; OR; SD; UT; VT; WA; WY; HI; BC; Mexico; Eurasia; Australia
[WildflowerSearch map]
from FNA
AZ; NV; TX
Discussion

In North America, Grimmia trichophylla is principally a lowland species, occurring in the mountains up to about 1000 m., rarely higher. In the Southern Hemisphere, it may be found up to 4000 m. In New Zealand, the species is common, and in contrast to G. trichophylla in North America, frequently bears capsules. The New Zealand plants are usually smaller than American specimens, and the leaves are frequently contorted. The nearly cosmopolitan G. trichophylla has many phenotypes, and numerous subspecies and varieties have been described. In damp and shaded habitats, the awns may be short, just as in dry unfavorable habitats at high altitudes, where stunted specimens may occur with small, short leaves and reduced awns, or even with muticous leaves. Grimmia trichophylla has frequently been confused with related species such as 36. G. muehlenbeckii and 34. G. lisae (see discussions thereunder for identification details). Robust forms of G. trichophylla have been mistaken for G. austrofunalis (H. C. Greven 1997, 2003), which does not occur in North America. Although some of those plants have leaves of equal length along the stem, characteristic of G. austrofunalis, they also have both leaf margins recurved, and the medial and outer basal laminal cells are longer and more robust than in that species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Grimmia americana is a rare endemic, currently known only from three sites. Until 1999, the species was known only from its type locality in Jeff Davis County of western Texas. A second locality was reported by J. Muñoz (1999) in Arizona and a third site by L. R. Stark et al. (2002) in Nevada. Thus, it is reasonable to expect G. americana to occur also in southern New Mexico. It is recognized as a member of subg. Grimmia by its immersed, ventricose capsule, with a short mitrate calyptra and eccentric seta attachment. The overall habit of the species also is similar to that of the other members of the subgenus. However, its basal laminal cells are thick-walled while other members have thin cell walls. H. A. Crum (1994c) implied that G. americana is similar to G. anodon, but the former is readily separated by its fully developed, perforated peristome, rostellate operculum, and thick-walled basal laminal cells. Grimmia crinitoleucophaea, also most commonly found in the American Southwest, is dioicous, has a rudimentary peristome, and thin-walled basal laminal cells.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 257. Treatment authors: Roxanne I. Hastings, Henk C. Greven. FNA vol. 27, p. 232. Treatment authors: Roxanne I. Hastings, Henk C. Greven.
Parent taxa Grimmiaceae > subfam. Grimmioideae > Grimmia > subg. Rhabdogrimmia Grimmiaceae > subfam. Grimmioideae > Grimmia > subg. Grimmia
Sibling taxa
G. alpestris, G. americana, G. anodon, G. anomala, G. arizonae, G. atrata, G. attenuata, G. brittoniae, G. caespiticia, G. crinitoleucophaea, G. donniana, G. elatior, G. elongata, G. funalis, G. hamulosa, G. hartmanii, G. incurva, G. laevigata, G. leibergii, G. lesherae, G. lisae, G. longirostris, G. mariniana, G. mollis, G. montana, G. moxleyi, G. muehlenbeckii, G. nevadensis, G. olneyi, G. orbicularis, G. ovalis, G. pilifera, G. plagiopodia, G. pulvinata, G. ramondii, G. reflexidens, G. serrana, G. sessitana, G. shastae, G. teretinervis, G. torquata, G. unicolor
G. alpestris, G. anodon, G. anomala, G. arizonae, G. atrata, G. attenuata, G. brittoniae, G. caespiticia, G. crinitoleucophaea, G. donniana, G. elatior, G. elongata, G. funalis, G. hamulosa, G. hartmanii, G. incurva, G. laevigata, G. leibergii, G. lesherae, G. lisae, G. longirostris, G. mariniana, G. mollis, G. montana, G. moxleyi, G. muehlenbeckii, G. nevadensis, G. olneyi, G. orbicularis, G. ovalis, G. pilifera, G. plagiopodia, G. pulvinata, G. ramondii, G. reflexidens, G. serrana, G. sessitana, G. shastae, G. teretinervis, G. torquata, G. trichophylla, G. unicolor
Name authority Greville: Fl. Edin., 235. (1824) E. B. Bartram: Bryologist 32: 8, fig. 1. (1929)
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