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grimmia dry rock moss, twisted grimmia

Habit Plants in soft, readily disintegrating cushions, yellow-green to brownish, occasionally light green, blackish to red-brown inside.
Stems

1–4 cm, small central strand present.

Gemmae

brown, multicellular, present on the abaxial side of distal leaves.

Leaves

contorted when dry, patent when moist, lanceolate, 1.5–2 × 0.3–0.5 mm, keeled, margins slightly recurved proximally, plane distally, awns very short and smooth, occasionally absent, costa weak proximally, projecting on abaxial side;

basal juxtacostal laminal cells linear, extremely sinuose, thick-walled;

basal marginal laminal cells long rectangular, thin-walled, hyaline;

medial laminal cells rectangular, extremely sinuose, thick-walled;

distal laminal cells 1-stratose.

Seta

slightly curved, straight when dry, 3–5 mm.

Sexual condition

dioicous.

Capsule

sporadically present, exserted, ovoid, brown, smooth, striate when dry and empty, exothecial cells thin-walled, annulus present, operculum rostrate, peristome teeth yellowish, short, split in distal part, weakly papillose.

Calyptra

mitrate.

Grimmia torquata

Habitat Damp, frequently vertical faces of acidic rock
Elevation moderate to high elevations (200-4000 m) (moderate to high elevations (700-13100 ft))
Distribution
from FNA
AK; AZ; CA; CO; ID; MT; NV; OR; WA; WY; HI; AB; BC; NL; NT; ON; QC; YT; Mexico; Eurasia; n Africa; Greenland
Discussion

Grimmia torquata is a montane-alpine species with a preference for shaded habitats. It grows in hemispherical cushions on steep, damp rock walls. Preferred substrates are granite, gneiss, quartzite, and schist. Its distribution reaches from near sea level in the Arctic to above 4000 m on Mexican and Hawaiian volcanoes. The species is easily recognized by its yellow-brown cushions of plants, which have contorted leaves when dry, and their brown gemmae, which are borne at the bases of older leaves. In densely shaded habitats, the cushions are light green; on exposed rock, they are usually brown. Although the species has a wide distribution, it is seldom present in quantity, usually growing in a small number of cushions in one habitat. Sporophytes are very rare worldwide and have not been found in eastern North America (H. A. Crum and L. E. Anderson 1981).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 256.
Parent taxa Grimmiaceae > subfam. Grimmioideae > Grimmia > subg. Rhabdogrimmia
Sibling taxa
G. alpestris, G. americana, G. anodon, G. anomala, G. arizonae, G. atrata, G. attenuata, G. brittoniae, G. caespiticia, G. crinitoleucophaea, G. donniana, G. elatior, G. elongata, G. funalis, G. hamulosa, G. hartmanii, G. incurva, G. laevigata, G. leibergii, G. lesherae, G. lisae, G. longirostris, G. mariniana, G. mollis, G. montana, G. moxleyi, G. muehlenbeckii, G. nevadensis, G. olneyi, G. orbicularis, G. ovalis, G. pilifera, G. plagiopodia, G. pulvinata, G. ramondii, G. reflexidens, G. serrana, G. sessitana, G. shastae, G. teretinervis, G. trichophylla, G. unicolor
Synonyms G. pellucida, G. prolifera, G. pseudotorquata, G. tortifolia
Name authority Drummond: Musc. Scot. 2: no. 28. (1825)
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