Grimmia ramondii |
Grimmia hartmanii |
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hartman's dry rock moss |
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Habit | Plants in robust, loose, arched-ascending, readily disintegrating patches, brownish green distally, blackish proximally. | Plants in ascending, dichotomously branched patches, green. |
Stems | 5–10 cm, central strand absent. |
2–8 cm, central strand absent. |
Gemmae | absent. |
in clusters, mulberry- or raspberry-shaped, brown, borne on leaf apices, usually present. |
Leaves | loosely appressed when dry, patent when moist, broadly ovate-lanceolate, tapering to an acute, slightly toothed apex, 2–3 × 0.4–0.6 mm, keeled, margins recurved on both sides, awns absent, costa stout, with two abaxial lamellae (or wings) forming parallel ridges along its length; basal juxtacostal laminal cells yellow, linear, sinuose, thick-walled; basal marginal laminal cells quadrate to short-rectangular, slightly thick-walled; medial laminal cells quadrate to rectangular, sinuous, thick-walled; distal laminal cells 1-stratose, margins 2-stratose. |
slightly contorted with often secund apices when dry, distal leaves usually falcate-secund when moist, broadly lanceolate, tapering to an acuminate apex, 2.5–4.5 × 0.4–0.6 mm, keeled, margins usually recurved on one side, awns very short, denticulate, costa firm, projecting on abaxial side; basal juxtacostal laminal cells short- to long-rectangular, slightly sinuose, thick-walled; basal marginal laminal cells quadrate, thickened transverse walls; medial laminal cells quadrate to short-rectangular, slightly sinuose, thick-walled; distal laminal cells 1-stratose, margins 2-stratose. |
Seta | cygneous to arcuate at maturity, flexuose when old, 3–5 mm. |
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Sexual condition | dioicous. |
dioicous. |
Capsule | occasionally present, exserted, obloid, yellowish green to yellowish brown, plicate when empty, exothecial cells thin-walled, annulus present, operculum rostrate, peristome teeth purple, divided nearly to base into two partly adhering segments, papillose. |
extremely rare, exserted, yellowish green, clavate-obloid, smooth, exothecial cells thick-walled, annulus present, operculum rostrate, peristome teeth orange, deeply split, perforated distally, papillose. |
Calyptra | mitrate. |
mitrate.] |
[seta straight to flexuose, 3–4 mm. | ||
Grimmia ramondii |
Grimmia hartmanii |
|
Habitat | Dry to damp acidic rock | Shaded boulders, especially granite, in woodlands |
Elevation | low to high elevations (0-2000 m) (low to high elevations (0-6600 ft)) | low to moderate elevations (100-1500 m) (low to moderate elevations (300-4900 ft)) |
Distribution |
AK; CA; ID; MI; MT; OR; WA; AB; BC; NF; NT; Greenland; Europe; Asia (China, Japan); n Africa |
UT; VT; PE; QC; Eurasia; n Africa |
Discussion | Grimmia ramondii occurs near sea level in arctic areas and along the West Coast. In the latter region, however, it is more common in mid-elevation montane sites and may also be found above the tree line. It differs from most Grimmia species by its lack of awns, winged costa, and tall, loose growth form. At first sight, G. ramondii has the habit of a Racomitrium. However, the sporophyte has an arcuate rather than straight seta, and the capsules are striate rather than smooth. Because of its habit and the absence of a central strand, K. G. Limpricht ([1885–]1890–1903, vol. 1) placed it, together with G. hartmanii and G. atrata, in Dryptodon, a genus intermediate between Grimmia and Racomitrium. There are many other Grimmia species, however, without a central strand, and G. hartmanii and G. atrata have already been removed from Dryptodon, so there is no reason to maintain that monotypic genus. Grimmia ramondii may be confused with G. hartmanii and G. elatior. However, the lack of awns, the Racomitrioideae-like areolation, and the distinct costal wings distinguish it easily from those species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Grimmia hartmanii is principally a lowland species, occurring in the mountains up to 1500 m. With the separation of G. anomala from it, G. hartmanii is much less common in North America than was previously thought. It is known principally from the Northeast. Although it is usually found in densely shaded habitats, in areas with high humidity it can establish itself in exposed sites. It more closely resembles some species of Racomitrioideae than it does species of its own genus, but it may be recognized by the usually spreading falcate-secund distal leaves, the very small awns, and the terminal clusters of brown gemmae, which are conspicuous and usually present. Capsules are extremely rare, and are unknown in North America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 256. | FNA vol. 27, p. 251. |
Parent taxa | Grimmiaceae > subfam. Grimmioideae > Grimmia > subg. Rhabdogrimmia | Grimmiaceae > subfam. Grimmioideae > Grimmia > subg. Rhabdogrimmia |
Sibling taxa | ||
Synonyms | Pterigynandrum ramondii, Dryptodon patens, G. curvata, Racomitrium patens | |
Name authority | (Lamarck & De Candolle) Margadant: Lindbergia 1: 128. (1972) | Schimper: Syn. Musc. Eur., 214. (1860) |
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