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Habit Plants 10–20 mm (–70 mm for G. atrata).
Stem

central strand present or absent.

Gemmae

absent.

Leaves

narrowly ovate-lanceolate to oblong-lanceolate, keeled, margins plane, recurved or incurved, costa prominent;

basal marginal cells hyaline or not;

distal lamina 2-stratose or with at least several marginal rows 2-stratose.

Seta

straight, centrally attached to capsule.

Sexual condition

autoicous or dioicous.

Capsule

immersed to exserted, smooth (except G. mariniana);

stomata present or absent, small, at neck to part way up capsule.

Calyptra

mitrate to cucullate, covering operculum.

Grimmia subg. Guembelia

Distribution
North America; Mexico; Central America; South America; Eurasia; Africa; Pacific Islands; Australia; Antarctic
Discussion

Species 41 (13 in the flora).

Members of subg. Guembelia are recognized by their thick, keeled leaves and usually long, straight setae with smooth capsules. Most occur on dry acidic rock, with three species growing on damp to wet acidic rock and one on dry calcareous rock.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 233.
Parent taxa Grimmiaceae > subfam. Grimmioideae > Grimmia
Subordinate taxa
Synonyms subg. Guembelia
Name authority (Hampe) Schimper: Coroll. Bryol. Eur., 48. (1856)
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