Gratiola quartermaniae
Gratiola lutea
limestone hedge-hyssop, Quarterman's hedge-hyssop
golden hedge-hyssop, gratiole dorée
erect, simple or few-branched, (7–)10–21(–30) cm, glabrous or glabrate proximally, sparsely glandular-puberulent distally.
decumbent to ascending or erect, simple or few-branched, 5–47 cm, glabrous proximally, sparsely glandular-puberulent distally.
blade linear to linear-lanceolate or lanceolate-elliptic, often falcate, (16–)18–30(–43) × (1–)2.5–5 mm, margins entire or with 1 or 2(or 3) pairs of blunt teeth distally, apex obtuse to acute, surfaces glabrous or glabrate.
blade lanceolate-ovate to oblong or ovate-elliptic, sometimes linear-lanceolate on submerged plants, 5–26 × (1.5–)3–7(–9) mm, margins entire or with 1 or 2 pairs of teeth, apex obtuse, rarely acute, surfaces glabrous.
slender, 7–14(–22) mm, length 0.5–1(–1.6) times bract, sparsely glandular-puberulent;
bracteoles 2, 2–4.5 mm.
slender, (3–)5–20 mm, length 0.4–2.1 times bract, sparsely glandular-puberulent;
bracteoles 2, 2–4 mm.
sepals distinct, lanceolate, 2.7–5 mm;
corolla 7–9 mm, tube and limb white tinged with pink or purple, veins white to greenish white or lavender;
style 3–4.6 mm.
sepals distinct, linear-lanceolate to lanceolate, 3.5–7 mm;
corolla 8–15 mm, tube and limb yellow, rarely white or cream, veins yellow to orangish yellow, rarely white;
style 3.5–5 mm.
ovoid, 3.4–5.1 × 3–4.5 mm.
ovoid, 2.2–4.5 × 2–3.5 mm.
0.4–0.7 mm.
0.4–0.6 mm.
= 28.
Gratiola quartermaniae
Gratiola lutea
Populations of Gratiola quartermaniae are concentrated in northern Alabama and central Tennessee. Disjunct populations are known from the Edwards Plateau of Texas, Will County, Illinois, and southeastern Ontario. D. Estes and R. L. Small (2007) discussed the distribution and ecology of G. quartermaniae.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
The name Gratiola aurea has been used widely for G. lutea due to questions about the applicability of the name by Rafinesque, which predates the name by Muhlenberg by two years. F. W. Pennell (1935) reviewed the nomenclatural history, noting that the type on which the name by Rafinesque is based, a collection by A. Michaux deposited at P, had been identified by A. Gray, M. L. Fernald, and S. F. Blake as G. neglecta. However, Pennell determined that the collection by Michaux was a mixed gathering and that the name by Rafinesque applies to the yellow-flowered element thereof.
Gratiola lutea (as G. aurea) has been reported from Crittenden County, Arkansas (W. H. Wilcox 1973); it is excluded from Arkansas in recent state checklists (E. B. Smith 1991; J. L. Gentry et al. 2013). Reports from Alabama may be based on misidentified specimens. A specimen identified as G. lutea has been collected in Japan (D. Estes 2008), where it was presumably introduced.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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