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golden hedge-hyssop, gratiole dorée

granite hedge-hyssop

Habit Perennials. Annuals.
Stems

decumbent to ascending or erect, simple or few-branched, 5–47 cm, glabrous proximally, sparsely glandular-puberulent distally.

erect, simple, sometimes few-branched, (7–)9–21(–29) cm, glabrous or glabrate proximally, glandular-puberulent distally.

Leaves

blade lanceolate-ovate to oblong or ovate-elliptic, sometimes linear-lanceolate on submerged plants, 5–26 × (1.5–)3–7(–9) mm, margins entire or with 1 or 2 pairs of teeth, apex obtuse, rarely acute, surfaces glabrous.

blade lanceolate-ovate to oblong, not falcate, 6–13(–18) × 1–3(–5) mm, margins entire or with 1 or 2(or 3) pairs of blunt teeth distally, apex obtuse, surfaces glabrate or glandular-puberulent.

Pedicels

slender, (3–)5–20 mm, length 0.4–2.1 times bract, sparsely glandular-puberulent;

bracteoles 2, 2–4 mm.

slender, (5–)8–17(–22) mm, length 0.9–2(–2.3) times bract, sparsely to densely glandular-puberulent;

bracteoles 2, 2–4.5 mm.

Flowers

sepals distinct, linear-lanceolate to lanceolate, 3.5–7 mm;

corolla 8–15 mm, tube and limb yellow, rarely white or cream, veins yellow to orangish yellow, rarely white;

style 3.5–5 mm.

sepals distinct, lanceolate, 2–4.5 mm;

corolla 7–9 mm, tube and limb white tinged with pink or purple, veins lavender or purple;

style 1.9–2.2 mm.

Capsules

ovoid, 2.2–4.5 × 2–3.5 mm.

subglobular to ovoid, 2.5–3.6 × 2–3.7 mm.

Seeds

0.4–0.6 mm.

0.3–0.5 mm.

2n

= 28.

Gratiola lutea

Gratiola graniticola

Phenology Flowering May–Oct. Flowering Apr–May.
Habitat Acidic freshwater pondshores, blackwater stream banks, cypress savannas, acidic wetlands, swamps. Seasonal pools on granite outcrops.
Elevation 0–300 m. (0–1000 ft.) 100–300 m. (300–1000 ft.)
Distribution
from FNA
CT; DC; DE; FL; GA; IL; MA; MD; ME; MI; NC; ND; NH; NJ; NY; PA; RI; SC; VA; VT; WI; NB; NF; NS; ON; QC [Introduced in e Asia (Japan)]
from FNA
GA; SC
[BONAP county map]
Discussion

The name Gratiola aurea has been used widely for G. lutea due to questions about the applicability of the name by Rafinesque, which predates the name by Muhlenberg by two years. F. W. Pennell (1935) reviewed the nomenclatural history, noting that the type on which the name by Rafinesque is based, a collection by A. Michaux deposited at P, had been identified by A. Gray, M. L. Fernald, and S. F. Blake as G. neglecta. However, Pennell determined that the collection by Michaux was a mixed gathering and that the name by Rafinesque applies to the yellow-flowered element thereof.

Gratiola lutea (as G. aurea) has been reported from Crittenden County, Arkansas (W. H. Wilcox 1973); it is excluded from Arkansas in recent state checklists (E. B. Smith 1991; J. L. Gentry et al. 2013). Reports from Alabama may be based on misidentified specimens. A specimen identified as G. lutea has been collected in Japan (D. Estes 2008), where it was presumably introduced.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Gratiola graniticola is known from about a dozen counties in the southeastern piedmont of northern Georgia and north-central South Carolina, where it occurs only on granite flatrocks (D. Estes and R. L. Small 2007; D. F. Brunton 2009).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 267. FNA vol. 17, p. 267.
Parent taxa Plantaginaceae > Gratiola Plantaginaceae > Gratiola
Sibling taxa
G. amphiantha, G. brevifolia, G. ebracteata, G. floridana, G. graniticola, G. heterosepala, G. neglecta, G. quartermaniae, G. ramosa, G. torreyi, G. virginiana, G. viscidula
G. amphiantha, G. brevifolia, G. ebracteata, G. floridana, G. heterosepala, G. lutea, G. neglecta, G. quartermaniae, G. ramosa, G. torreyi, G. virginiana, G. viscidula
Synonyms G. aurea
Name authority Rafinesque: Med. Repos., hexade 3, 2: 333. (1811) D. Estes: J. Bot. Res. Inst. Texas 1: 166, figs. 3A,D, 9. (2007)
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