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Boggs Lake hedge-hyssop

pool sprite, snorkelwort

Habit Annuals. Annuals.
Stems

ascending to erect, simple or few-branched, 2–12 cm, glabrous proximally, glabrous or glandular-puberulent distally.

erect, simple or few-branched, (7–)9–21(–29) cm, glabrous or glabrate proximally, glabrate or glandular-puberulent distally.

Leaves

blade linear-lanceolate to oblanceolate, elliptic, or oblong, 4–8(–20) × 1.5–3.5 mm, margins entire, apex obtuse to rounded, surfaces glabrous.

dimorphic, submersed basal and clustered, floating paired at ends of branches to 70 mm, blade of submersed lanceolate to oblanceolate or oblong, 1–7 × 0.5–3 mm, margins entire, apex obtuse to acute, surfaces glabrous, blade of floating elliptic to ovate or nearly round, 3–10 × 2–9 mm, margins entire, apex obtuse to rounded or retuse, surfaces glabrous.

Pedicels

slender, 2–14(–28) mm, length 0.9–2.3 times bract, sparsely to densely glandular-puberulent;

bracteoles 0.

stout, 0.1–3 mm, length less than 0.1 times bract, glabrous;

bracteoles 0.

Flowers

sepals connate proximally, calyx distinctly bilaterally symmetric, lobes elliptic-oblanceolate, 3.5–6 mm;

corolla 4–7.5(–9) mm, tube yellow or greenish yellow, veins purple, limb white abaxially, greenish yellow adaxially;

style 1.5–2 mm.

sepals connate proximally, calyx lobes obovate to oblong, 0.7–1 mm;

corolla 3–4 mm, tube and limb white tinged pink or purple, veins lavender or purple;

style 0.5–1 mm.

Capsules

subglobular to ovoid, 3.5–4.7 × 3–4.5 mm.

bilaterally symmetric, obcordiform, 2–3 × 3–4 mm.

Seeds

0.5–0.7 mm.

(0.3–)0.4–0.5 mm.

2n

= 18.

Gratiola heterosepala

Gratiola amphiantha

Phenology Flowering Apr–Sep. Flowering Mar–Apr.
Habitat Shallow water and exposed mud of vernal pools, wet meadows, lake margins. Shallow, ephemeral pools on exposed granite outcrops.
Elevation 0–1900 m. (0–6200 ft.) 100–300 m. (300–1000 ft.)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; GA; SC
[BONAP county map]
Discussion

Phylogenetic studies confirmed that Gratiola amphiantha, long treated in monospecific Amphianthus, is embedded in Gratiola (D. Estes and R. L. Small 2008). Gratiola amphiantha is listed as a federally-threatened species by the United States Fish and Wildlife Service.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 267. FNA vol. 17, p. 266.
Parent taxa Plantaginaceae > Gratiola Plantaginaceae > Gratiola
Sibling taxa
G. amphiantha, G. brevifolia, G. ebracteata, G. floridana, G. graniticola, G. lutea, G. neglecta, G. quartermaniae, G. ramosa, G. torreyi, G. virginiana, G. viscidula
G. brevifolia, G. ebracteata, G. floridana, G. graniticola, G. heterosepala, G. lutea, G. neglecta, G. quartermaniae, G. ramosa, G. torreyi, G. virginiana, G. viscidula
Synonyms Amphianthus pusillus
Name authority H. Mason & Bacigalupi: Madroño 12: 150, figs. 1–8. (1954) D. Estes & R. L. Small: Syst. Bot. 33: 181. (2008)
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