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pool sprite, snorkelwort

American hedge-hyssop, clammy hedge-hyssop, common American hedge hyssop, gratiole négligée

Habit Annuals. Annuals.
Stems

erect, simple or few-branched, (7–)9–21(–29) cm, glabrous or glabrate proximally, glabrate or glandular-puberulent distally.

ascending or erect, few- to much-branched, rarely simple, (4–)10–40 cm, glabrous or glandular-puberulent proximally, glandular-puberulent distally.

Leaves

dimorphic, submersed basal and clustered, floating paired at ends of branches to 70 mm, blade of submersed lanceolate to oblanceolate or oblong, 1–7 × 0.5–3 mm, margins entire, apex obtuse to acute, surfaces glabrous, blade of floating elliptic to ovate or nearly round, 3–10 × 2–9 mm, margins entire, apex obtuse to rounded or retuse, surfaces glabrous.

blade linear to narrowly elliptic, oblanceolate, or elliptic-obovate, rarely falcate, 10–65 × 3–11(–18) mm, margins with (1 or)2–5(–7) pairs of blunt to sharp teeth, apex acute, surfaces glabrate or glandular-puberulent.

Pedicels

stout, 0.1–3 mm, length less than 0.1 times bract, glabrous;

bracteoles 0.

slender, 8–35 mm, length (0.3–)0.4–0.9(–1.3) times bract, sparsely to densely glandular-puberulent;

bracteoles 2, 2.5–7 mm.

Flowers

sepals connate proximally, calyx lobes obovate to oblong, 0.7–1 mm;

corolla 3–4 mm, tube and limb white tinged pink or purple, veins lavender or purple;

style 0.5–1 mm.

sepals distinct, lanceolate to narrowly lanceolate-elliptic, 2.5–6 mm;

corolla 7–12 mm, tube yellow, yellowish white, or yellowish green, veins brownish violet, limb white, sometimes tinged lavender;

style 3–4 mm.

Capsules

bilaterally symmetric, obcordiform, 2–3 × 3–4 mm.

ovoid, 2.6–6 × 3–5 mm.

Seeds

(0.3–)0.4–0.5 mm.

0.4–0.7 mm.

2n

= 18.

= 14, 16.

Gratiola amphiantha

Gratiola neglecta

Phenology Flowering Mar–Apr. Flowering Mar–Oct.
Habitat Shallow, ephemeral pools on exposed granite outcrops. Wet meadows, stream banks, shorelines of ponds, mudflats, salt marshes, crop fields.
Elevation 100–300 m. (300–1000 ft.) 0–2400 m. (0–7900 ft.)
Distribution
from FNA
AL; GA; SC
[BONAP county map]
from FNA
AL; AR; AZ; CA; CO; CT; DC; DE; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NS; ON; QC; SK [Introduced in Europe (Finland, France)]
[WildflowerSearch map]
[BONAP county map]
Discussion

Phylogenetic studies confirmed that Gratiola amphiantha, long treated in monospecific Amphianthus, is embedded in Gratiola (D. Estes and R. L. Small 2008). Gratiola amphiantha is listed as a federally-threatened species by the United States Fish and Wildlife Service.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Gratiola neglecta exhibits the widest range and broadest ecological amplitude of the North American Gratiola species. Besides occurring in a variety of freshwater wetland communities, it sometimes is found in salt marshes and damp or wet agricultural fields.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 266. FNA vol. 17, p. 268.
Parent taxa Plantaginaceae > Gratiola Plantaginaceae > Gratiola
Sibling taxa
G. brevifolia, G. ebracteata, G. floridana, G. graniticola, G. heterosepala, G. lutea, G. neglecta, G. quartermaniae, G. ramosa, G. torreyi, G. virginiana, G. viscidula
G. amphiantha, G. brevifolia, G. ebracteata, G. floridana, G. graniticola, G. heterosepala, G. lutea, G. quartermaniae, G. ramosa, G. torreyi, G. virginiana, G. viscidula
Synonyms Amphianthus pusillus G. neglecta var. glaberrima
Name authority D. Estes & R. L. Small: Syst. Bot. 33: 181. (2008) Torrey: Cat. Pl. New York, 89. (1819)
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