Gratiola amphiantha |
Gratiola |
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pool sprite, snorkelwort |
hedge-hyssop, hyssop |
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Habit | Annuals. | Herbs, annual or perennial. | ||||||||||||||||||||||||||||||||||||||||||||||||
Stems | erect, simple or few-branched, (7–)9–21(–29) cm, glabrous or glabrate proximally, glabrate or glandular-puberulent distally. |
decumbent to ascending or erect, glabrous or glandular-hairy. |
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Leaves | dimorphic, submersed basal and clustered, floating paired at ends of branches to 70 mm, blade of submersed lanceolate to oblanceolate or oblong, 1–7 × 0.5–3 mm, margins entire, apex obtuse to acute, surfaces glabrous, blade of floating elliptic to ovate or nearly round, 3–10 × 2–9 mm, margins entire, apex obtuse to rounded or retuse, surfaces glabrous. |
cauline (also basal and clustered in G. amphiantha), opposite, monomorphic (dimorphic in G. amphiantha); petiole absent; blade not fleshy, not leathery, margins entire or toothed, surfaces often glandular-punctate. |
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Inflorescences | axillary, flowers 1 or 2 per node; bracts present. |
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Pedicels | stout, 0.1–3 mm, length less than 0.1 times bract, glabrous; bracteoles 0. |
present, sometimes short; bracteoles present or absent, sepal-like. |
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Flowers | sepals connate proximally, calyx lobes obovate to oblong, 0.7–1 mm; corolla 3–4 mm, tube and limb white tinged pink or purple, veins lavender or purple; style 0.5–1 mm. |
bisexual; sepals 5, distinct or proximally connate, calyx ± bilaterally symmetric, campanulate, lobes (sepals when distinct) linear to lanceolate, obovate, or oblong; corolla white, cream, or yellow, sometimes pinkish, tube yellowish, brownish, or purplish, often with dark veins, bilaterally symmetric, bilabiate, tubular, tube base not spurred or gibbous, lobes 5, abaxial 3, adaxial 2, nearly completely connate and appearing as 1 emarginate lobe, throat glabrous or pubescent; stamens 2, medially adnate to corolla, filaments glabrous, pollen sacs perpendicular to filaments, connective dilated; staminodes 0 or 2, minute; ovary 2-locular, placentation axile; stigma capitate or lamelliform. |
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Fruits | capsules, dehiscence septicidal, sometimes also loculicidal. |
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Capsules | bilaterally symmetric, obcordiform, 2–3 × 3–4 mm. |
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Seeds | (0.3–)0.4–0.5 mm. |
ca. 200, yellow to brown or black, oblong to obpyramidal or short-cylindric, wings absent. |
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× = 9. | ||||||||||||||||||||||||||||||||||||||||||||||||||
2n | = 18. |
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Gratiola amphiantha |
Gratiola |
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Phenology | Flowering Mar–Apr. | |||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Shallow, ephemeral pools on exposed granite outcrops. | |||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 100–300 m. (300–1000 ft.) | |||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
AL; GA; SC |
North America; Mexico; South America; Central America (Guatemala); Europe; e Asia (Japan); Africa (Morocco); Pacific Islands (New Zealand); Australia |
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Discussion | Phylogenetic studies confirmed that Gratiola amphiantha, long treated in monospecific Amphianthus, is embedded in Gratiola (D. Estes and R. L. Small 2008). Gratiola amphiantha is listed as a federally-threatened species by the United States Fish and Wildlife Service. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 30 (13 in the flora). Members of Gratiola sometimes are confused with Lindernia and Mecardonia. Sophronanthe often is included in Gratiola; molecular and morphological data support separating the two as sister genera (D. Estes 2008). This treatment was prepared in part from information contained in a partial manuscript submitted by D. Estes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 266. | FNA vol. 17, p. 264. | ||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Amphianthus pusillus | Amphianthus | ||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | D. Estes & R. L. Small: Syst. Bot. 33: 181. (2008) | Linnaeus: Sp. Pl. 1: 17. (1753): Gen. Pl. ed. 5, 11. (1754) | ||||||||||||||||||||||||||||||||||||||||||||||||
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