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fowl manna grass, glycerie striee, ridged glyceria, ridged manna grass

Habit Plants perennial. Plants usually perennial, sometimes annual; cespitose, sometimes rhizomatous.
Culms

20-80 (100) cm tall, (1.5)2-3.5 mm thick, not or only slightly spongy, sometimes rooting at the lower nodes.

annual, not woody, not branching above the base;

internodes hollow.

Sheaths

smooth to scabridulous, keeled, sometimes weakly so;

ligules 1-4 mm, usually rounded, sometimes acute to mucronate, erose-lacerate;

blades 12-30 cm long, 2-6 mm wide, abaxial surfaces smooth or scabridulous, adaxial surfaces scabridulous to scabrous.

closed for their whole length or almost so;

collars without tufts of hair on the sides;

auricles sometimes present;

ligules hyaline, glabrous, often lacerate, occasionally ciliate, those of the lower and upper cauline leaves usually similar;

pseudopetioles absent;

blades linear to narrowly lanceolate, venation parallel, cross venation sometimes evident;

cross sections non-Kranz, without arm or fusoid cells;

epidermes without microhairs, sometimes papillate.

Panicles

6-25 cm long, 2.5-21 cm wide, pyramidal, open, nodding;

branches 5-13 cm, straight to lax, lower branches usually strongly divergent to drooping at maturity, sometimes ascending, with 15-50 spikelets, these often confined to the distal 2/3;

pedicels 0.5-7 mm.

Inflorescences

terminal panicles or racemes;

disarticulation above the glumes and beneath the florets or below the glumes.

Spikelets

1.8-4 mm long, 1.2-2.9 mm wide, laterally compressed, oval in side view, with 3-7 florets.

2.5-60 mm, not viviparous, slightly to strongly laterally compressed, with 1-30 florets, proximal florets bisexual, distal 1-3 florets usually sterile, sometimes pistillate, sometimes reduced and amalgamated into a knob- or club-shaped rudiment;

rachillas prolonged beyond the base of the distal floret.

Glumes

ovate, 1-1.5 times longer than wide, narrowing from midlength or above, veins terminating below the apical margins, apices often splitting with age;

lower glumes 0.5-1.2 mm, rounded to obtuse;

upper glumes 0.6-1.2 mm, acute or rounded;

rachilla internodes 0.1-0.6 mm;

lemmas 1.2-2 mm, ovate in dorsal view, veins raised, scabridulous over and between the veins, apices acute, prow-shaped;

paleas slightly shorter than to equaling the lemmas, lengths 1.5-3 times widths, keeled, keels not winged, tips pointing towards each other, apices narrowly notched between the keels;

anthers 2, (0.2)0.4-0.6 mm, purple or yellow.

exceeded by the distal florets, shorter than to longer than the adjacent lemmas, mostly membranous, scarious distally, 1-11-veined, apices usually rounded to acute;

florets laterally or dorsally compressed;

calluses blunt, glabrous or with hairs;

lemmas of sexual florets rectangular or ovate, mostly membranous, scarious distally, often with a purplish band adjacent to the scarious apices, (4)5-15-veined, veins not converging distally, often prominent, unawned or awned, awns not branched, apices entire to bilobed or bifid, awns straight, subterminal or from the sinuses;

paleas from shorter than to longer than the lemmas, similar in texture, 2-veined, veins keeled, sometimes winged;

lodicules 2, fleshy, usually connate into a single structure, without a membranous wing, truncate, not ciliate, not or scarcely veined;

anthers 1, 2, or 3;

ovaries glabrous;

styles 2-branched, bases persistent, branches plumose distally.

Caryopses

0.5-2 mm.

ovoid to ellipsoid, longitudinally grooved or not;

hila usually linear;

embryos less than 1/3 as long as the caryopses.

x

= (8)9, 10.

2n

= 20 [reports of 28 are questionable].

Glyceria striata

Poaceae tribe Meliceae

Distribution
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Glyceria striata grows in bogs, along lakes and streams, and in other wet places. Its range extends from Alaska to Newfoundland and south into Mexico. Plants from the eastern portion of the range have sometimes been treated as G. striata var. striata, and those from the west as G. striata var. stricta (Scribn.) Fernald. Eastern plants tend to have somewhat narrower leaves and thinner culms than western plants, but the variation appears continuous. In the west, larger specimens are easy to confuse with G. elata. The two species are sometimes found growing together without hybridizing; this and molecular data (Whipple et al. [in prep.]) support their recognition as separate species. The differences between the two in growth habit and stature are evident in the field; they are not always evident on herbarium specimens. In its overall aspect, G. striata also resembles G. pulchella, but it has somewhat more lax panicle branches in addition to smaller spikelets and florets.

Glyceria xgatineauensis Bowden is a sterile hybrid between G. striata and G. melicaria. It resembles G. melicaria but has longer (up to 12 cm), less appressed panicle branches and is a triploid with 2n = 30. It was described from a population near Eardley, Quebec. An additional specimen, tentatively identified as G. xgatineauensis, was collected in 1929 from French Creek in Upshur County, West Virginia.

Glyceria xottawensis Bowden is a sterile hybrid between G. striata and G. canadensis. It is intermediate between the two parents, and is known only from the original populations near Ottawa. It has sometimes been included in G. xlaxa (Scribn.) Scribn. [=G. canadensis var. laxa]; that taxon often produces viable seed, indicating that it is not a hybrid.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

There are approximately 130 species and 8 or 9 genera in the Meliceae. Four of the genera are monotypic. Melica and Glyceria, the two largest genera, are well represented in North America. Pleuropogon and Schizachne are primarily North American, but extend into eastern Asia.

Molecular studies (e.g., Soreng and Davis 2000; Grass Phylogeny Working Group 2001) show the tribe to be monophyletic and somewhat basal within the Pooideae. Members of the tribe are most easily recognized by the combination of closed leaf sheaths, scarious lemma apices, and non-converging lemma veins. The tribe also differs from other tribes in the Pooideae in having 2 unwinged lodicules that are usually connate into a single structure, and a base chromosome number of 9 or 10. Catabrosa and Briza, whose inclusion in the tribe was suggested by the preliminary results of Mejia-Saules and Bisby (2000), have more membranous lemma margins and free, winged lodicules. Briza also has open leaf sheaths and more convergent lemma veins. Their inclusion is not supported by the molecular data.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Calluses hairy; lemmas awned, awns 8-15 mm long, twisted, divergent to slightly geniculate
Schizachne
1. Calluses glabrous; lemmas unawned or awned, awns to 12 mm long, straight.
→ 2
2. Inflorescences racemes; palea keels winged, the wings notched and awned
Pleuropogon
2. Inflorescences usually panicles, racemes in depauperate specimens; palea keels not winged or the wings entire and unawned.
→ 3
3. Lower glumes 1-veined, 0.3-4.5 mm long; disarticulation always above the glumes; lemmas unawned, never with hairs more than 1 mm long; culms never with cormous bases; distal florets in the spikelets sometimes reduced, not forming a morphologically distinct rudiment; plants of wet meadows and streamsides
Glyceria
3. Lower glumes 1-9-veined, 2-16 mm long; disarticulation above or below the glumes; lemmas sometimes awned, sometimes with hairs longer than 1 mm; culms sometimes with cormous bases; distal florets in the spikelets often forming a morphologically distinct rudiment; plants of drier or well drained habitats
Melica
Source FNA vol. 24, p. 77. FNA vol. 24, p. 67. Author: Mary E. Barkworth;.
Parent taxa Poaceae > subfam. Pooideae > tribe Meliceae > Glyceria > sect. Striatae Poaceae > subfam. Pooideae
Sibling taxa
G. acutiflora, G. alnasteretum, G. borealis, G. canadensis, G. declinata, G. elata, G. fluitans, G. grandis, G. leptostachya, G. maxima, G. melicaria, G. notata, G. nubigena, G. obtusa, G. pulchella, G. septentrionalis, G. ×occidentalis
Subordinate taxa
Glyceria, Melica, Pleuropogon, Schizachne
Synonyms G. striata var. stricta, G. striata subsp. stricta, Panicularia nervata, G. nervata
Name authority (Lam.) Hitchc. Endl.
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