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great Smoky Mountain glyceria, great Smoky Mountain mannagrass

American glyceria, American manna grass, manna grass, reed mannagrass, tall mannagrass

Habit Plants perennial. Plants perennial.
Culms

100-200 cm tall, 3-5 mm thick, smooth.

50-150 (200) cm tall, 8-12 mm thick, erect or decumbent and rooting at the base.

Sheaths

smooth or scabridulous, weakly keeled;

ligules 1-1.5 mm, truncate;

blades to 45 cm long, 6-10 mm wide, abaxial surfaces smooth or scabrous, adaxial surfaces scabrous.

smooth or scabridulous, keeled;

ligules 1-5 (7) mm, truncate to rounded, ligules of the lower leaves stiff at the base, ligules of the upper leaves flexible throughout;

blades 25-43 cm long, 4.5-15 mm wide.

Panicles

20-30 cm long, 7.5-14 cm wide, open, pyramidal;

branches 7.5-14 cm, spreading or reflexed, lax, with 16-80 spikelets;

pedicels 2-7 mm.

16-42 cm long, 12-20 cm wide, open;

branches (7)10-18 cm, lax, widely divergent to drooping, with 35-80+ spikelets;

pedicels 1-15 mm.

Spikelets

3.5-5.5 mm long, 2-3(3.5) mm wide, laterally compressed, oval in side view, with 3-5 florets.

3.2-10 mm long, 2-3 mm wide, somewhat laterally compressed, oval to elliptic in side view, with 4-10 florets.

Glumes

tapering from below midlength to the narrowly (< 45°) acute apices, veins not extending to the apices;

lower glumes 0.8-1.5 mm;

upper glumes 1.8-2.2 mm;

rachilla internodes about 0.5 mm;

lemmas 2.2-2.7 mm, 0.9-1.1 mm wide in dorsal view, veins distinctly raised, usually smooth over and between the veins, sometimes scabridulous over the veins, apices acute, prow-shaped;

paleas slightly shorter than the lemmas, lengths 2-2.7 times widths, keels not winged, tips incurved, apices narrowly notched between the keels;

anthers 2, about 1.5 mm, dehiscent at maturity.

mostly hyaline, usually the midvein of 1 or both glumes extending to the apices, apices acute;

lower glumes 1-2.3 mm;

upper glumes 1.5-2.7 mm;

rachilla internodes 0.5-0.8 mm;

lemmas 1.8-3 mm, prominently (5)7-veined, veins often scabridulous, intercostal regions smooth, apices rounded to truncate, sometimes erose, almost flat at maturity;

paleas from shorter than to slightly longer than the lemmas, lengths more than 3 times widths, keels not winged, ciliolate, tips not strongly incurved, truncate to notched between the keels;

anthers 3, 0.5-1.2 mm.

Caryopses

about 1.5 mm.

1-1.5 mm.

2n

= 40.

=20.

Glyceria nubigena

Glyceria grandis

Distribution
from FNA
NC; TN
[BONAP county map]
from FNA
AK; AZ; CA; CO; CT; DE; IA; ID; IL; IN; MA; MD; ME; MI; MN; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; LB; MB; NB; NS; NT; ON; PE; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Glyceria nubigena is known only from moist areas of balds and high ridges in the Great Smoky Mountains of North Carolina and Tennessee.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Glyceria grandis grows on banks and in the water of streams, ditches, ponds, and wet meadows, from Alaska to Newfoundland and south in the mountains to California, Arizona, and New Mexico in the western United States, and to Virginia and Tennessee in the eastern United States. It is similar to G. maxima, differing primarily in its shorter, flatter lemmas and shorter anthers. It is also confused with G. elata and Torreyochloa pallida. It differs from the former in having acute glumes with long veins, more evenly dark florets, flatter lemma apices, and paleal keel tips that do not point towards each other. It differs from Torreyochloa pallida in its closed leaf sheaths and 1-veined glumes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Spikelets 3.2-6.4 mm long, with 4-8 florets
var. grandis
1. Spikelets 6-10 mm long, with 5-10 florets
var. komarovii
Source FNA vol. 24, p. 75. FNA vol. 24, p. 71.
Parent taxa Poaceae > subfam. Pooideae > tribe Meliceae > Glyceria > sect. Striatae Poaceae > subfam. Pooideae > tribe Meliceae > Glyceria > sect. Hydropoa
Sibling taxa
G. acutiflora, G. alnasteretum, G. borealis, G. canadensis, G. declinata, G. elata, G. fluitans, G. grandis, G. leptostachya, G. maxima, G. melicaria, G. notata, G. obtusa, G. pulchella, G. septentrionalis, G. striata, G. ×occidentalis
G. acutiflora, G. alnasteretum, G. borealis, G. canadensis, G. declinata, G. elata, G. fluitans, G. leptostachya, G. maxima, G. melicaria, G. notata, G. nubigena, G. obtusa, G. pulchella, G. septentrionalis, G. striata, G. ×occidentalis
Subordinate taxa
G. grandis var. grandis, G. grandis var. komarovii
Synonyms G. maxima subsp. grandis, G. grandis forma pallescens
Name authority W.A. Anderson S. Watson
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