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northwestern manna grass, western manna grass

fowl manna grass, glycerie striee, ridged glyceria, ridged manna grass

Habit Plants perennial. Plants perennial.
Culms

60-160 cm tall, 2.5-5 mm thick, erect or decumbent and rooting from the lower nodes.

20-80 (100) cm tall, (1.5)2-3.5 mm thick, not or only slightly spongy, sometimes rooting at the lower nodes.

Sheaths

smooth to scabridulous, keeled, sometimes weakly so;

ligules 7-12 mm;

blades 20-30 cm long, (2.5)4-12 mm wide, adaxial surfaces scabridulous, occasionally papillose.

smooth to scabridulous, keeled, sometimes weakly so;

ligules 1-4 mm, usually rounded, sometimes acute to mucronate, erose-lacerate;

blades 12-30 cm long, 2-6 mm wide, abaxial surfaces smooth or scabridulous, adaxial surfaces scabridulous to scabrous.

Panicles

20-50 cm long, 2-15 cm wide, usually narrow, open at anthesis;

branches 4.5-18 cm, somewhat lax, usually ascending, strongly divergent at anthesis, with 2-8 spikelets, pedicels 1.5-8 mm.

6-25 cm long, 2.5-21 cm wide, pyramidal, open, nodding;

branches 5-13 cm, straight to lax, lower branches usually strongly divergent to drooping at maturity, sometimes ascending, with 15-50 spikelets, these often confined to the distal 2/3;

pedicels 0.5-7 mm.

Spikelets

13-23 mm long, 1.5-3.5 mm wide, cylindrical and terete, except at anthesis when slightly laterally compressed, rectangular in side view, with 6-13 florets.

1.8-4 mm long, 1.2-2.9 mm wide, laterally compressed, oval in side view, with 3-7 florets.

Glumes

acute to obtuse;

lower glumes 1.1-2.8 mm;

upper glumes 2.9-3.7 mm, about twice as long as the lower glumes;

rachilla internodes 1-2.8 mm;

lemmas 4.5-5.9 mm, scabridulous, midveins extending to within 0.1 mm of the apical margins, apices acute, usually slightly lobed or irregularly crenate;

paleas usually shorter than or equaling the lemmas, sometimes slightly longer, keels winged, apices shallowly notched to slightly bifid, teeth to 0.2 mm, parallel;

anthers 2, 0.6-1.6 mm.

ovate, 1-1.5 times longer than wide, narrowing from midlength or above, veins terminating below the apical margins, apices often splitting with age;

lower glumes 0.5-1.2 mm, rounded to obtuse;

upper glumes 0.6-1.2 mm, acute or rounded;

rachilla internodes 0.1-0.6 mm;

lemmas 1.2-2 mm, ovate in dorsal view, veins raised, scabridulous over and between the veins, apices acute, prow-shaped;

paleas slightly shorter than to equaling the lemmas, lengths 1.5-3 times widths, keeled, keels not winged, tips pointing towards each other, apices narrowly notched between the keels;

anthers 2, (0.2)0.4-0.6 mm, purple or yellow.

Caryopses

0.5-2 mm.

2n

= 40.

= 20 [reports of 28 are questionable].

Glyceria ×occidentalis

Glyceria striata

Distribution
from FNA
CA; ID; NV; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Glyceria ×occidentalis has hitherto been considered an uncommon native species that grows along lakes, ponds, and streams, and in marshy areas of western North America. It differs from other species in the region primarily in its longer lemmas and anthers. Studies of chloroplast DNA in western North American species of Glyceria demonstrated that, contrary to C.L. Hitchcock's (1969) conclusion, G. fluitans is present in western North America, and that all specimens being identified as G. ×occidentalis had cpDNA resembling that of G. leptostachya or G. fluitans; there was no distinctive G. ×occidentalis cpDNA (Whipple et al. [in press]). This strongly suggests that G. ×occidentalis is a series of reciprocal hybids, and probably backcrosses, between G. fluitans and G. leptostachya. As the key indicates, G. ×occidentalis is intermediate between its two putative parents. The cpDNA study also confirmed that G. declinata is distinct from G. ×occidentalis (see discussion under that species).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Glyceria striata grows in bogs, along lakes and streams, and in other wet places. Its range extends from Alaska to Newfoundland and south into Mexico. Plants from the eastern portion of the range have sometimes been treated as G. striata var. striata, and those from the west as G. striata var. stricta (Scribn.) Fernald. Eastern plants tend to have somewhat narrower leaves and thinner culms than western plants, but the variation appears continuous. In the west, larger specimens are easy to confuse with G. elata. The two species are sometimes found growing together without hybridizing; this and molecular data (Whipple et al. [in prep.]) support their recognition as separate species. The differences between the two in growth habit and stature are evident in the field; they are not always evident on herbarium specimens. In its overall aspect, G. striata also resembles G. pulchella, but it has somewhat more lax panicle branches in addition to smaller spikelets and florets.

Glyceria xgatineauensis Bowden is a sterile hybrid between G. striata and G. melicaria. It resembles G. melicaria but has longer (up to 12 cm), less appressed panicle branches and is a triploid with 2n = 30. It was described from a population near Eardley, Quebec. An additional specimen, tentatively identified as G. xgatineauensis, was collected in 1929 from French Creek in Upshur County, West Virginia.

Glyceria xottawensis Bowden is a sterile hybrid between G. striata and G. canadensis. It is intermediate between the two parents, and is known only from the original populations near Ottawa. It has sometimes been included in G. xlaxa (Scribn.) Scribn. [=G. canadensis var. laxa]; that taxon often produces viable seed, indicating that it is not a hybrid.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 85. FNA vol. 24, p. 77.
Parent taxa Poaceae > subfam. Pooideae > tribe Meliceae > Glyceria > sect. Glyceria Poaceae > subfam. Pooideae > tribe Meliceae > Glyceria > sect. Striatae
Sibling taxa
G. acutiflora, G. alnasteretum, G. borealis, G. canadensis, G. declinata, G. elata, G. fluitans, G. grandis, G. leptostachya, G. maxima, G. melicaria, G. notata, G. nubigena, G. obtusa, G. pulchella, G. septentrionalis, G. striata
G. acutiflora, G. alnasteretum, G. borealis, G. canadensis, G. declinata, G. elata, G. fluitans, G. grandis, G. leptostachya, G. maxima, G. melicaria, G. notata, G. nubigena, G. obtusa, G. pulchella, G. septentrionalis, G. ×occidentalis
Synonyms G. striata var. stricta, G. striata subsp. stricta, Panicularia nervata, G. nervata
Name authority (Piper) J.C. Nelson (Lam.) Hitchc.
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