Glyceria ×occidentalis |
Glyceria grandis |
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northwestern manna grass, western manna grass |
American glyceria, American manna grass, manna grass, reed mannagrass, tall mannagrass |
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Habit | Plants perennial. | Plants perennial. | ||||
Culms | 60-160 cm tall, 2.5-5 mm thick, erect or decumbent and rooting from the lower nodes. |
50-150 (200) cm tall, 8-12 mm thick, erect or decumbent and rooting at the base. |
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Sheaths | smooth to scabridulous, keeled, sometimes weakly so; ligules 7-12 mm; blades 20-30 cm long, (2.5)4-12 mm wide, adaxial surfaces scabridulous, occasionally papillose. |
smooth or scabridulous, keeled; ligules 1-5 (7) mm, truncate to rounded, ligules of the lower leaves stiff at the base, ligules of the upper leaves flexible throughout; blades 25-43 cm long, 4.5-15 mm wide. |
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Panicles | 20-50 cm long, 2-15 cm wide, usually narrow, open at anthesis; branches 4.5-18 cm, somewhat lax, usually ascending, strongly divergent at anthesis, with 2-8 spikelets, pedicels 1.5-8 mm. |
16-42 cm long, 12-20 cm wide, open; branches (7)10-18 cm, lax, widely divergent to drooping, with 35-80+ spikelets; pedicels 1-15 mm. |
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Spikelets | 13-23 mm long, 1.5-3.5 mm wide, cylindrical and terete, except at anthesis when slightly laterally compressed, rectangular in side view, with 6-13 florets. |
3.2-10 mm long, 2-3 mm wide, somewhat laterally compressed, oval to elliptic in side view, with 4-10 florets. |
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Glumes | acute to obtuse; lower glumes 1.1-2.8 mm; upper glumes 2.9-3.7 mm, about twice as long as the lower glumes; rachilla internodes 1-2.8 mm; lemmas 4.5-5.9 mm, scabridulous, midveins extending to within 0.1 mm of the apical margins, apices acute, usually slightly lobed or irregularly crenate; paleas usually shorter than or equaling the lemmas, sometimes slightly longer, keels winged, apices shallowly notched to slightly bifid, teeth to 0.2 mm, parallel; anthers 2, 0.6-1.6 mm. |
mostly hyaline, usually the midvein of 1 or both glumes extending to the apices, apices acute; lower glumes 1-2.3 mm; upper glumes 1.5-2.7 mm; rachilla internodes 0.5-0.8 mm; lemmas 1.8-3 mm, prominently (5)7-veined, veins often scabridulous, intercostal regions smooth, apices rounded to truncate, sometimes erose, almost flat at maturity; paleas from shorter than to slightly longer than the lemmas, lengths more than 3 times widths, keels not winged, ciliolate, tips not strongly incurved, truncate to notched between the keels; anthers 3, 0.5-1.2 mm. |
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Caryopses | 1-1.5 mm. |
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2n | = 40. |
=20. |
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Glyceria ×occidentalis |
Glyceria grandis |
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Distribution |
CA; ID; NV; OR; WA; BC
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AK; AZ; CA; CO; CT; DE; IA; ID; IL; IN; MA; MD; ME; MI; MN; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; AB; BC; LB; MB; NB; NS; NT; ON; PE; QC; SK; YT
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Discussion | Glyceria ×occidentalis has hitherto been considered an uncommon native species that grows along lakes, ponds, and streams, and in marshy areas of western North America. It differs from other species in the region primarily in its longer lemmas and anthers. Studies of chloroplast DNA in western North American species of Glyceria demonstrated that, contrary to C.L. Hitchcock's (1969) conclusion, G. fluitans is present in western North America, and that all specimens being identified as G. ×occidentalis had cpDNA resembling that of G. leptostachya or G. fluitans; there was no distinctive G. ×occidentalis cpDNA (Whipple et al. [in press]). This strongly suggests that G. ×occidentalis is a series of reciprocal hybids, and probably backcrosses, between G. fluitans and G. leptostachya. As the key indicates, G. ×occidentalis is intermediate between its two putative parents. The cpDNA study also confirmed that G. declinata is distinct from G. ×occidentalis (see discussion under that species). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Glyceria grandis grows on banks and in the water of streams, ditches, ponds, and wet meadows, from Alaska to Newfoundland and south in the mountains to California, Arizona, and New Mexico in the western United States, and to Virginia and Tennessee in the eastern United States. It is similar to G. maxima, differing primarily in its shorter, flatter lemmas and shorter anthers. It is also confused with G. elata and Torreyochloa pallida. It differs from the former in having acute glumes with long veins, more evenly dark florets, flatter lemma apices, and paleal keel tips that do not point towards each other. It differs from Torreyochloa pallida in its closed leaf sheaths and 1-veined glumes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 85. | FNA vol. 24, p. 71. | ||||
Parent taxa | ||||||
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Subordinate taxa | ||||||
Synonyms | G. maxima subsp. grandis, G. grandis forma pallescens | |||||
Name authority | (Piper) J.C. Nelson | S. Watson | ||||
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