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northwestern manna grass, western manna grass

Canada mannagrass, Canadian mannagrass, rattlesnake manna grass, rattlesnake-grass

Habit Plants perennial. Plants perennial.
Culms

60-160 cm tall, 2.5-5 mm thick, erect or decumbent and rooting from the lower nodes.

60-150 cm tall, 2.5-5 mm thick, erect or the bases decumbent.

Sheaths

smooth to scabridulous, keeled, sometimes weakly so;

ligules 7-12 mm;

blades 20-30 cm long, (2.5)4-12 mm wide, adaxial surfaces scabridulous, occasionally papillose.

retrorsely scabridulous to scabrous, keeled;

ligules 2-6 mm;

blades 8-36 cm long, 3-8 mm wide, abaxial surfaces smooth or scabrous, adaxial surfaces scabridulous to scabrous.

Panicles

20-50 cm long, 2-15 cm wide, usually narrow, open at anthesis;

branches 4.5-18 cm, somewhat lax, usually ascending, strongly divergent at anthesis, with 2-8 spikelets, pedicels 1.5-8 mm.

10-30 cm long, 10-20 cm wide, pyramidal, open, nodding;

branches 7-20 cm, lax, divergent, often drooping, with 15-60+ spikelets;

pedicels 2.5-9 mm.

Spikelets

13-23 mm long, 1.5-3.5 mm wide, cylindrical and terete, except at anthesis when slightly laterally compressed, rectangular in side view, with 6-13 florets.

3-8 mm long, (2.5)3-5 mm wide, laterally compressed, oval in side view, with 2-10 florets.

Glumes

acute to obtuse;

lower glumes 1.1-2.8 mm;

upper glumes 2.9-3.7 mm, about twice as long as the lower glumes;

rachilla internodes 1-2.8 mm;

lemmas 4.5-5.9 mm, scabridulous, midveins extending to within 0.1 mm of the apical margins, apices acute, usually slightly lobed or irregularly crenate;

paleas usually shorter than or equaling the lemmas, sometimes slightly longer, keels winged, apices shallowly notched to slightly bifid, teeth to 0.2 mm, parallel;

anthers 2, 0.6-1.6 mm.

narrowing from midlength or above to the broadly (> 45°) acute or rounded apices, 1-veined, veins terminating below the apices;

lower glumes 0.6-2.4 mm, ovate to rectangular;

upper glumes 1.5-2.5 mm, lanceolate;

rachilla internodes 0.2-0.5 mm;

lemmas 1.8-4 mm, ovate in dorsal view, 5-7-veined, veins evident but not raised distally, smooth over and between the veins, apices acute, prow-shaped;

paleas 0.1-0.8 mm shorter than lemmas, lengths 1.5-1.8 times widths, almost round in dorsal view, keels well developed, not winged, tips incurved, apices narrowly notched between the keels;

anthers 2, 0.4-0.5 mm, dehiscent at maturity.

Caryopses

1.5-2 mm.

2n

= 40.

Glyceria ×occidentalis

Glyceria canadensis

Distribution
from FNA
CA; ID; NV; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
CT; DC; DE; IL; IN; MA; MD; ME; MI; MN; NC; NH; NJ; NY; OH; OR; PA; RI; VA; VT; WA; WI; WV; BC; MB; NB; NL; NS; ON; PE; QC; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Glyceria ×occidentalis has hitherto been considered an uncommon native species that grows along lakes, ponds, and streams, and in marshy areas of western North America. It differs from other species in the region primarily in its longer lemmas and anthers. Studies of chloroplast DNA in western North American species of Glyceria demonstrated that, contrary to C.L. Hitchcock's (1969) conclusion, G. fluitans is present in western North America, and that all specimens being identified as G. ×occidentalis had cpDNA resembling that of G. leptostachya or G. fluitans; there was no distinctive G. ×occidentalis cpDNA (Whipple et al. [in press]). This strongly suggests that G. ×occidentalis is a series of reciprocal hybids, and probably backcrosses, between G. fluitans and G. leptostachya. As the key indicates, G. ×occidentalis is intermediate between its two putative parents. The cpDNA study also confirmed that G. declinata is distinct from G. ×occidentalis (see discussion under that species).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Glyceria canadensis is an attractive native species that grows in swamps, bogs, lakeshore marshes, and wet woods throughout much of eastern North America, extending from eastern Saskatchewan to Newfoundland, Illinois, and northeastern Tennessee. It is now established in western North America, having been introduced as a weed in cranberry farms. It forms sterile hybrids with G. striata; the hybrids are called G. xottawensis Bowden. For further comments, see p. 77.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Lemmas 2.4-4 mm long; spikelets 5-8 mm long, with 4-10 florets; lower glumes 1.6-2.4 mm long; upper glumes acute
var. canadensis
1. Lemmas 1.8-2.5 mm long; spikelets 3-5 mm long, with 2-5 florets; lower glumes 0.6-1.3 mm long; upper glumes usually rounded, sometimes acute
var. laxa
Source FNA vol. 24, p. 85. FNA vol. 24, p. 79.
Parent taxa Poaceae > subfam. Pooideae > tribe Meliceae > Glyceria > sect. Glyceria Poaceae > subfam. Pooideae > tribe Meliceae > Glyceria > sect. Striatae
Sibling taxa
G. acutiflora, G. alnasteretum, G. borealis, G. canadensis, G. declinata, G. elata, G. fluitans, G. grandis, G. leptostachya, G. maxima, G. melicaria, G. notata, G. nubigena, G. obtusa, G. pulchella, G. septentrionalis, G. striata
G. acutiflora, G. alnasteretum, G. borealis, G. declinata, G. elata, G. fluitans, G. grandis, G. leptostachya, G. maxima, G. melicaria, G. notata, G. nubigena, G. obtusa, G. pulchella, G. septentrionalis, G. striata, G. ×occidentalis
Subordinate taxa
G. canadensis var. canadensis, G. canadensis var. laxa
Synonyms Poa canadensis
Name authority (Piper) J.C. Nelson (Michx.) Trin.
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