Flora of North America species comparison

Varieties 2 (2 in the flora).

Characters useful in recognizing specimens of Geum macrophyllum are yellow petals, epicalyx bractlets often absent, proximal style segment sparsely to densely stipitate-glandular, and fruiting receptacles puberulent. Across its broad North American range from Alaska to California and Nova Scotia, G. macrophyllum exhibits considerable variation. Based largely on the shape of the terminal leaflet of the basal leaves and the degree of dissection and shape of the divisions of the distal cauline leaves, P. A. Rydberg (1913b) distinguished three species within the range of variation treated here as one species. Basal leaves of G. macrophyllum in the strict sense (var. macrophyllum in this treatment) have relatively large reniform to rounded terminal leaflets and the distal cauline leaves are three-cleft into rhombic or cuneate lobes. The basal-leaf terminal leaflets of G. perincisum (= var. perincisum) are only slightly larger than the laterals and are deeply lobed into rhombic-obovate segments; the distal cauline leaves are dissected into oblanceolate divisions. Both basal and cauline leaves of G. oregonense (= var. perincisum) are intermediate between those of G. macrophyllum and G. perincisum. W. Gajewski (1955) crossed all three taxa under discussion and examined leaf morphology and cytology of the F1 and F2 hybrids. He concluded that they were distinct but not yet completely separated species. Fairly well correlated with the more dissected leaves of G. oregonense and G. perincisum is the presence of minute stalked glands on the pedicels. The treatment here follows H. M. Raup (1931) in recognizing two varieties of G. macrophyllum based more on pedicel glandularity than leaf morphology.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

At the beginning of the twentieth century, E. L. Greene described over a dozen species belonging to the Geum triflorum complex based on differences in leaf form and indument, the relative length and shape of the epicalyx bractlets and sepals, and petal length and shape. Most of these species were quickly reduced to synonymy by other botanists. When specimens are examined from across the continent, most of the characters used to separate species in the G. triflorum complex show nearly continuous variation. It seems best to treat these variants as belonging to one species. Whether and how to classify the variation within the species will remain controversial. Some character expressions correlate reasonably well with dividing the species into two varieties, as was first proposed by N. C. Fassett (1928). Variety triflorum occurs east of the Rocky Mountains and is typical of the grasslands covering the northern Great Plains; it also is the variety found in the mountains of Arizona and New Mexico. Variety ciliatum is common throughout the rest of the Rocky Mountains, the Sierra Nevada, and Cascade Range. Intermediate specimens occur here and there, particularly in Alberta and British Columbia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)