Geum canadense |
Geum geniculatum |
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benoîte du Canada, white avens |
bent avens |
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Habit | Plants leafy-stemmed. | Plants leafy-stemmed. |
Stems | 30–100 cm, glabrate to downy, hairs to 1.5 mm, sometimes glandular. |
50–84 cm, sparsely hirsute proximally, hirsute distally. |
Leaves | basal 10–25 cm, blade simple or pinnate, major leaflets 3–5, plus 0–4 minor basal ones, terminal leaflet larger than major laterals; cauline 3–8 cm, stipules ± free, 4–13 × 1–7 mm, blade 3-foliolate or simple and 3-lobed to unlobed. |
basal 14–30 cm, blade interruptedly lyrate-pinnate, major leaflets 1–5, mixed with 2–5 minor ones, terminal leaflet slightly to much larger than major laterals; cauline 4–12 cm, stipules ± free, 7–20 × 4–8 mm, blade pinnate, 3-foliolate, or simple and 3-lobed. |
Inflorescences | 3–15-flowered. |
5–15-flowered. |
Pedicels | densely hairy, hairs of varying lengths, few long stiff ones, sometimes glandular. |
densely pilose, some hairs septate-glandular. |
Flowers | erect; epicalyx bractlets 0.5–1.5 mm; hypanthium green; sepals spreading but soon reflexed, 3–6 mm; petals spreading, white, obovate to oblong, (3–)4–8 mm, ± equal to or slightly longer than sepals, apex rounded. |
nodding; epicalyx bractlets 2.5–3 mm; hypanthium green with purple tinge at base to greenish maroon; sepals erect to erect-spreading, 5–8 mm; petals erect, white, pink, or lavender, spatulate-obovate, 5–7 mm, ± equal to sepals, apex rounded, truncate, or shallowly emarginate. |
Fruiting tori | sessile, densely bristly, hairs 1–2.3 mm. |
sessile, densely bristly. |
Fruiting styles | geniculate-jointed, proximal segment persistent, 2–8 mm, apex hooked, usually glabrous, sometimes sparsely hairy or stipitate-glandular, distal segment deciduous, 1–2 mm, pilose in basal 1/2, hairs much longer than diam. of style. |
geniculate-jointed, proximal segment persistent, 2–4.5 mm, apex hooked, septate-glandular hairs on basal 1/2, distal segment very tardily deciduous, 5–8 mm, pilose except apical 1–2 mm, with some glandular hairs. |
2n | = 42. |
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Geum canadense |
Geum geniculatum |
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Phenology | Flowering spring–summer. | Flowering summer. |
Habitat | Lowlands and upland forests, meadows, along streams, thickets, bottomland hardwoods, swamps | Mountain balds, seeps, stream banks, cliffs, wooded boulder fields |
Elevation | 0–600 m (0–2000 ft) | 1600–1900 m (5200–6200 ft) |
Distribution |
AL; AR; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; WY; MB; NB; NS; ON; PE; QC; Mexico (Chiapas)
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NC; TN |
Discussion | Varieties and forms have been described in an effort to classify the variation encompassed in Geum canadense. In the eastern half of the United States, it is by far the most common, widespread, and variable of the Geum species. Nearly all writers of recent floras have not found it worthwhile to apply names to the variants. Perhaps the most distinctive and worthy of further consideration are plants from the southwestern corner of the range in Arkansas, Oklahoma, and Texas. These plants, some of which fit Fernald and Weatherby’s description of var. texanum, bloom from late March through May, significantly earlier than the rest of the species, which typically flowers after June first, even in the other southern states. Geum canadense hybridizes with G. urbanum (= G. ×catlingii J.-P. Bernard & R. Gauthier); see discussion under 15. G. urbanum. Geum album J. F. Gmelin is a superfluous name that pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Geum geniculatum is enigmatic and is currently known from 15 populations in western North Carolina and eastern Tennessee. As a rare plant, it has attracted far less attention than G. radiatum, which is also known from the same region. Although G. geniculatum is not listed federally as endangered or threatened, it is listed as threatened by North Carolina and as endangered by Tennessee. W. Gajewski (1957) speculated that Geum geniculatum is a hybrid, with one of the parents being G. rivale. K. R. Robertson (1974) pointed out that the nearest locality for G. rivale is in West Virginia and that pollen stainability of G. geniculatum is very high. Geum geniculatum may be of hybrid origin; it cannot be a first-generation hybrid. North American taxonomists have treated it as a valid species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 68. | FNA vol. 9, p. 65. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Colurieae > Geum | Rosaceae > subfam. Rosoideae > tribe Colurieae > Geum |
Sibling taxa | ||
Synonyms | G. camporum, G. canadense var. brevipes, G. canadense var. camporum, G. canadense var. grimesii, G. canadense var. texanum | |
Name authority | Jacquin: Hort. Bot. Vindob. 2: 82, plate 175. 1772–1773 | Michaux: Fl. Bor.-Amer. 1: 300. (1803) |
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