Gentiana newberryi |
Gentianaceae |
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alpine gentian, Newberry's gentian |
gentian family |
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Habit | Herbs perennial, 0.1–1.5(–3.5) dm (below flowers), glabrous. | Herbs [shrubs, trees], annual, biennial, or perennial, autotrophic, with green stems and leaves, or mycotrophic; when strongly mycotrophic, stems and leaves weakly chlorophyllous (only in Bartonia) or yellowish, whitish, purplish, or buff, or lacking chlorophyll (only in Voyria). | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | 1–5, arising laterally below rosettes, from a stout tap root, tufted, decumbent. |
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Leaves | basal and cauline; blades of basal rosette and proximal cauline leaf blades widely spatulate to obovate or oblanceolate, 0.8–5 cm × 2–25 mm, apex obtuse or mucronate, at least these leaves with blades less than 6 times as long as wide, distal cauline leaves few, with blades oblanceolate to lanceolate or linear, 2–5 cm × 2–5 mm, apices acute. |
cauline, often also basal, opposite, whorled, or rarely alternate, sessile or petiolate, simple; stipules absent [rarely present as ocreae]; blade margins entire. |
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Inflorescences | terminal, flowers usually solitary, occasionally 2 or 3. |
cymes (sometimes racemoid, spicoid, or capitate), thyrses, or verticillasters, or solitary flowers; flowers pedicellate or sessile. |
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Flowers | calyx 14–30 mm, lobes linear to narrowly ovate, (4–)6–12 mm, margins not ciliate; corolla white or blue, campanulate, open, 23–55 mm, lobes spreading, elliptic-obovate, 7–17 mm, free portions of plicae divided into 2 triangular, serrate to lacerate segments; anthers distinct. |
bisexual or occasionally some unisexual [all unisexual on some or all plants], homostylous [heterostylous], protandrous and outbreeding or less often homogamous and autogamous, radially [somewhat bilaterally] symmetric, 4–12(–14)-merous [rarely 3-, 6-, or 16-merous] except for carpels; perianth hypogynous, calyx and usually corolla persistent; calyx green or occasionally ± hyaline (absent in Obolaria), sepals connate or some [or all] nearly distinct, lobes imbricate in bud, often ± unequal, colleters often present adaxially near base; corolla petaloid, petals connate, lobes contorted in bud or rarely imbricate (Obolaria, Voyria), spurs present only in Halenia, 1 per petal; stamens epipetalous, isomerous and alternate with petals, all fertile [rarely some sterile], equal [unequal]; filaments free or connected by a corona; anthers 2-locular, dehiscing longitudinally [with terminal pores], remaining straight, recurving, or coiling helically or circinately, distinct or (only in some spp. |
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Fruits | capsular, dehiscence septicidal or rarely rupturing irregularly (Obolaria) [indehiscent capsules, berries]. |
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Seeds | winged. |
few–very many, usually sessile; endosperm abundant and embryo small in autotrophic species, endosperm scant and embryo undifferentiated in completely mycotrophic species (Voyria). |
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Of | Gentiana) coherent; pistil 1, 2-carpellate; ovary 1[or 2]-locular; placentae 2, parietal [axile]; style present or absent, erect or initially deflexed to one side [declinate], uncleft, shallowly 2-cleft, or deeply cleft (Sabatia); stigmas 1 or 2, coiling only in Sabatia, decurrent on ovary (only in Lomatogonium, sometimes slightly so in Bartonia). |
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Gentiana newberryi |
Gentianaceae |
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Distribution |
w United States
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nearly worldwide |
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Discussion | Varieties 2 (2 in the flora). The two varieties of Gentiana newberryi intergrade extensively. The most distinctive form of var. newberryi, with relatively tall stems and medium to deep blue corollas, occurs in the northern part of the range of the species, from the Klamath and White mountains of California north into Oregon. Plants most clearly referable to var. tiogana prevail in the southern part of the range of the species, from Butte County south to Inyo and Tulare counties, California. In the central part of the range of the species, plant size and corolla color are less consistently correlated, with occasional plants combining low stature with deep blue corollas or tall stems with predominantly white or pale blue corollas. In that part of the range, corolla color may be highly variable within a single population. The leaves of Gentiana newberryi are thick-textured and distinctively concave, usually spoon-shaped, when fresh. Narrower leaves sometimes occur in var. tiogana, but many plants of that variety have widely spatulate leaves like those of var. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera ca. 100, species ca. 1800 (18 genera, 112 species in the flora). In the tribal classification by L. Struwe et al. (2002), genera 1–7 in this flora are in tribe Chironieae Dumortier, subtribe Chironiinae G. Don. Species of Chironieae generally lack nectaries, although Sabatia reportedly has indistinct nectaries at the base of the ovary. Genera 8–17 are in tribe Gentianeae Dumortier. Gentiana, in which the nectaries are on the gynophore, is in subtribe Gentianinae G. Don; the remaining genera of the Gentianeae in the flora area, all of which have epipetalous nectaries, are in subtribe Swertiinae Grisebach. Voyria constitutes the monogeneric tribe Voyrieae Gilg, in which the nectaries (when present) are on the ovary or the gynophore. Pedicel lengths given here refer to the true pedicels, between the most distal pair of bracts or bractlets and the calyx. In some genera, notably Centaurium, Sabatia, and Zeltnera, a flower terminating the ultimate branch of an inflorescence, directly subtended by bractlets, although sessile by this definition, may appear pedicellate. Corolla lengths as given are from the receptacle to the apices of the lobes (or plicae in Gentiana andrewsii). The Gentianaceae include many species esteemed in ornamental horticulture. In addition to those noted under the respective genera, the more important species in North American horticulture include Exacum affine Balfour f. ex Regel, Persian-violet, native to the island of Socotra, Yemen, which is widely grown as a florists’ pot plant. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 14. | FNA vol. 14. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Name authority | A. Gray: Proc. Amer. Acad. Arts 11: 84. (1876) | Jussieu | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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