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bottle gentian, closed gentian, gentiane close, meadow bottle gentian

King gentian, King's gentian, King's scepter gentian, King's-scepter or staff or Pacific gentian, staff gentian

Habit Herbs perennial, 2–8 dm, glabrous. Herbs perennial, 1–9 dm, glabrous.
Stems

1–10, terminal from caudex, erect or decumbent.

1–4(–10), terminal from caudex, decumbent to erect.

Leaves

cauline, ± evenly spaced;

blade ovate, 3–15 cm × 10–45 mm, apex acuminate.

cauline, gradually more widely spaced distally;

blade lanceolate to ovate or elliptic, 1–9 cm × 5–15(–20) mm, apex obtuse to acute.

Inflorescences

1–20-flowered heads, sometimes with additional flowers at 1–3 nodes, rarely on short branches.

solitary flowers or 2–5-flowered heads or umbellate cymes, often also on peduncles to 8 cm or in cymules on branches from distal 1–4(–7) nodes.

Flowers

calyx 8–22 mm, lobes spreading nearly horizontally, widely obovate or elliptic to orbiculate, 2–6(–10) mm, margins ciliate;

corolla blue or occasionally violet or white, tubular, completely closed, 23–40 mm, lobes incurved, ovate-triangular to semicircular, 0.7–2 mm, free portions of plicae ± as long and as wide as lobes, oblong, deeply and unequally bifid, summit erose;

anthers connate.

calyx 13–27 mm, tube rarely deeply cleft, lobes lanceolate to elliptic-ovate or rarely foliaceous, 6–15 mm, margins not ciliate;

corolla blue or rarely rose-violet or white, narrowly campanulate, opening only slightly with lobes ascending to incurved, 25–50 mm, lobes oblong-ovate to orbiculate, 5–10 mm, summit of plicae forming ± truncate, entire sinus, not or scarcely extending beyond bases of lobes;

anthers distinct.

Seeds

winged.

winged toward ends only.

2n

 = 26.

 = 26.

Gentiana clausa

Gentiana sceptrum

Phenology Flowering late summer–fall. Flowering summer–fall.
Habitat Moist, open woods, stream banks, roadsides, acid soils. Bogs, wet meadows.
Elevation 0–800 m. (0–2600 ft.) 0–1300 m. (0–4300 ft.)
Distribution
from FNA
CT; DC; MA; MD; ME; NC; NH; NJ; NY; OH; PA; RI; TN; VA; VT; WV; QC; restricted to higher elevations southward
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

The corollas of Gentiana andrewsii, G. austromontana, and G. clausa all remain completely and tightly closed but are pollinated by bumblebees, which force open the corollas. The fresh corollas of G. clausa are rounded at the summit, with the plicae concealed by the true lobes. In contrast, the fresh corollas of G. andrewsii and G. austromontana are more acute, with the plicae forming much or all of the visible summit. The corolla lobes of G. clausa are about as long and as wide as the free portions of the plicae, whereas those of G. andrewsii and G. austromontana are distinctly narrower than the plicae. Also, in contrast to those of other species of Gentiana in the flora area except for G. flavida and G. latidens, the calyx lobes of G. clausa when fresh spread almost horizontally rather than being nearly erect.

Reports of Gentiana clausa from Indiana to Missouri and elsewhere west of the range given here have been based on specimens of G. andrewsii var. dakotica, second- or later-generation plants derived from G. andrewsii × G. puberulenta, or other hybrids and introgressants. In these plants, in contrast to G. clausa, the sepals are lanceolate and nearly erect, and the lobes of the intact corolla do not entirely conceal the plicae.

Gentiana clausa is largely isolated ecologically and geographically, but a few hybrids with G. andrewsii, G. austromontana, G. decora, and G. saponaria are known.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plants of Gentiana sceptrum with ascending, narrowly lanceolate leaves conspicuously exceeded by the distal internodes have been called G. menziesii. Plants with spreading, elliptic to ovate, more closely spaced leaves have been called G. sceptrum var. cascadensis, although that variant might more appropriately be considered nomenclaturally typical. Studies for this flora have indicated that the extremes are connected by many intermediates, and that little correlation exists between leaf shape and geographic distribution, or between leaf shape and the other traits by which the segregates have been characterized, such as stature, erectness of stems, or numbers and sizes of flowers. Even within a relatively limited area, such as Vancouver Island, British Columbia, or Humboldt County, California, plants can be found with leaves ranging from narrowly lanceolate to widely elliptic.

Small plants with strongly decumbent stems, found at a few localities at or near the coast in California and southern Oregon, have been called Gentiana sceptrum var. humilis. Spathaceous calyces occasionally occur in these plants. Such plants are not recognized taxonomically here because larger plants approaching typical G. sceptrum have been found at the same localities or nearby, but they should be given further study.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 14. FNA vol. 14.
Parent taxa Gentianaceae > Gentiana Gentianaceae > Gentiana
Sibling taxa
G. affinis, G. algida, G. andrewsii, G. austromontana, G. autumnalis, G. calycosa, G. catesbaei, G. decora, G. douglasiana, G. flavida, G. fremontii, G. glauca, G. latidens, G. linearis, G. newberryi, G. nivalis, G. parryi, G. pennelliana, G. platypetala, G. plurisetosa, G. prostrata, G. puberulenta, G. rubricaulis, G. saponaria, G. sceptrum, G. setigera, G. villosa
G. affinis, G. algida, G. andrewsii, G. austromontana, G. autumnalis, G. calycosa, G. catesbaei, G. clausa, G. decora, G. douglasiana, G. flavida, G. fremontii, G. glauca, G. latidens, G. linearis, G. newberryi, G. nivalis, G. parryi, G. pennelliana, G. platypetala, G. plurisetosa, G. prostrata, G. puberulenta, G. rubricaulis, G. saponaria, G. setigera, G. villosa
Synonyms G. menziesii, G. sceptrum var. cascadensis, G. sceptrum var. humilis
Name authority Rafinesque: Med. Fl. 1: 210. (1828) Grisebach in W. J. Hooker: Fl. Bor.-Amer. 2: 57, plate 145. (1837)
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