Gentiana andrewsii |
Gentiana glauca |
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Andrew's bottle gentian, bottle or fringe bottle or prairie closed gentian, closed bottle gentian |
glaucous gentian, glaucous or blue-green or inky gentian, pale gentian |
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Habit | Herbs perennial, 1–12 dm, glabrous or rarely puberulent. | Herbs perennial, 0.2–1.6 dm, glabrous. | ||||
Stems | 1–20, terminal from caudex, decumbent to erect. |
erect, arising singly at intervals from slender, elongating, horizontal rhizomes, forming patches. |
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Leaves | cauline, ± evenly spaced; blade elliptic-oblong to lanceolate or narrowly ovate, 3–16 cm × 10–50 mm, apex acuminate. |
basal and cauline; cauline ± abruptly more widely spaced; blade elliptic to spatulate-obovate, apex obtuse; basal and rosette leaf blades 0.8–2(–2.7) cm × 4–12 mm; cauline blades 0.5–1.7 cm × 3–8(–12) mm. |
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Inflorescences | 1–25-flowered heads, often with additional flowers at 1–6(–9) nodes or on short branches. |
reduced, ± dense cymes, 2–5(–7)-flowered, also often a pair at most of the distal node, rarely solitary flowers. |
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Flowers | calyx 9–29 mm, lobes lanceolate to ovate or occasionally oblanceolate, 2–15 mm, margins ciliate; corolla blue, white, or rarely rose-violet, tubular, completely closed, 28–45 mm, lobes reduced to a mucro or ± triangular, 0.5–2(–3) mm, free portions of plicae oblong, shallowly and nearly symmetrically bifid, summit truncate, erose; anthers connate. |
calyx 5–7 mm, lobes ascending, lanceolate-triangular, 2–4 mm, margins not ciliate; corolla greenish blue or greenish yellow, rarely white, tubular, opening narrowly, (8–)12–20 mm, lobes ascending, triangular, 1.8–4 mm, free portions of plicae rounded, minutely erose; anthers distinct. |
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Seeds | winged. |
winged. |
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2n | = 24. |
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Gentiana andrewsii |
Gentiana glauca |
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Phenology | Flowering summer. | |||||
Habitat | Asia (n Japan, Kamchatka, coastal Siberia).. | |||||
Elevation | 0–2500 m. (0–8200 ft.) | |||||
Distribution |
North America
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AK; MT; WA; AB; BC; NT; YT; restricted to high elevations southward; Asia (n Japan, Kamchatka, coastal Siberia)
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Discussion | Varieties 2 (2 in the flora). Gentiana andrewsii is the only species of Gentiana in which the plicae of the corolla are distinctly longer than the lobes. Gentiana andrewsii has often been reported outside its actual range. Although the epithets of some of the species that have been confused with or considered inseparable from G. andrewsii have priority, the familiar name G. andrewsii is often misapplied, sometimes because it is assumed that any “closed gentian” is G. andrewsii. As G. clausa was not distinguished from G. andrewsii in standard floras prior to 1950, reports from the northeastern United States based on specimens identified before 1950 should be considered doubtful if the specimens have not been reexamined. Old reports from the southern Appalachians are also questionable because G. austromontana was not recognized until 1964. Some reports from the southeastern and south-central United States and along the Atlantic seaboard have been based on specimens of G. saponaria. True G. andrewsii is distinguishable as the only Gentiana species in which the corolla plicae distinctly exceed the minute lobes. The fringed tip of the completely closed corolla, at first white, soon turning reddish brown, is an excellent field mark for distinguishing G. andrewsii from G. clausa. In G. clausa, the summit of the intact corolla appears completely blue (in the typical color form), and the plicae are concealed. Gentiana andrewsii grows in calcareous soils and G. clausa in noncalcareous soils. Because of this ecological separation, there are only a few records of hybridization between Gentiana andrewsii and G. clausa. In the tall-grass prairies, G. andrewsii hybridizes with G. flavida, producing G. × pallidocyanea J. S. Pringle, and with G. puberulenta, producing G. × billingtonii Farwell (as species). Northward, it occasionally hybridizes with G. rubricaulis, producing G. × grandilacustris J. S. Pringle, and in the southeastern part of its range it hybridizes with G. saponaria. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The description by Kellogg of var. paulensis does not indicate clearly how this variety was believed to differ from Gentiana glauca elsewhere in its range. Specimens from the Pribilof Islands examined in studies for this flora, including an isotype and other plants from Saint Paul Island, do not appear to be taxonomically separable. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 14. | FNA vol. 14. | ||||
Parent taxa | Gentianaceae > Gentiana | Gentianaceae > Gentiana | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Dasystephana andrewsii, Pneumonanthe andrewsii | G. glauca var. paulensis | ||||
Name authority | Grisebach in W. J. Hooker: Fl. Bor.-Amer. 2: 55. (1837) | Pallas: Fl. Ross. 1(2): 104, plate 93, fig. 2. (1789) | ||||
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