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blanket-flower, fire wheel, gaillardia

lanceleaf blanketflower, lanceleaf gaillardia, Winkler's blanketflower

Habit Annuals, perennials, or subshrubs, 10–80+ cm (sometimes rhizomatous). Perennials (sometimes flowering first year) 10–60+ cm (sometimes rhizomatous).
Stems

usually erect, usually branched (from bases or throughout).

Leaves

basal, basal and cauline, or mostly cauline; alternate;

petiolate or sessile;

blades mostly elliptic, lanceolate, linear, oblong, obovate, ovate, or spatulate, often pinnately lobed, ultimate margins toothed or entire, faces usually scabrellous to villous (hairs simple or jointed), rarely glabrous, usually notably gland-dotted (except in 10. G. suavis).

cauline;

petiolar bases 0–3+ cm;

blades narrowly elliptic, linear, obovate, or spatulate, 15–60 × 3–12(–22) mm, (bases of distal often clasping) margins remotely toothed or entire, faces usually closely scabrellous (hairs sometimes crisped).

Peduncles

1–10(–20) cm.

Involucres

hemispheric to nearly rotate, mostly 9–20+ mm diam.

Receptacles

convex to hemispheric, smooth or pitted (often with scattered, hard, setiform enations), epaleate.

Ray florets

0, or 5–15+, usually neuter, rarely pistillate and fertile or styliferous and sterile;

corollas yellow to white or orange to red, purplish, or brown, often bicolored (sometimes “ray” corollas funnelform, 5-lobed, heads radiant).

usually 6–12(–15), sometimes 0;

corollas pinkish to purple or yellow to cream or white, 13–25 mm.

Disc florets

20–100+, bisexual, fertile;

corollas yellow or orange to red, purplish, or brown (distally ± shaggily hairy, hairs jointed), tubes much shorter than ampliate, campanulate to cylindric or urceolate throats, lobes 5, broadly deltate or ovate (± equal, sometimes attenuate or attenuate-terete apically; style-branch apices ± attenuate, usually hispidulous).

20–60(–100+);

corollas yellow or purple to purple-brown or bicolored, tubes 0.5–1 mm, throats stoutly cylindric to campanulate or urceolate, 3.5–5 mm, lobes attenuate-terete, 1.5–3 mm, jointed hairs to 0.3 mm.

Phyllaries

persistent, 14–40+ in 2–3+ series (reflexed in fruit, usually distinct, oblanceolate, lanceolate, narrowly triangular, or linear, subequal to unequal, usually chartaceous proximally, distally herbaceous, apices often attenuate).

15–26, lance-ovate to lanceolate, 6–14+ mm, scabrellous.

Heads

usually radiate or discoid (sometimes radiant), borne singly.

Cypselae

obpyramidal to clavate, ± 4-angled (lengths 1–1.5+ diams.), sparsely to densely hairy (at least proximally, hairs straight);

pappi usually persistent, of 6–10+ medially thickened, laterally scarious scales in 1–2 series (all, some, or none aristate), rarely 0.

1.5–2 mm, hairs 1 mm, inserted at bases and on angles and faces;

pappi of 8–10 lanceolate, aristate scales 5–7 mm (scarious bases 2–3.5 × 0.6–1 mm).

Receptacular

setae 0 or 0.1–0.5+ mm.

x

= 19.

2n

= 34, 68.

Gaillardia

Gaillardia aestivalis

Phenology Flowering Jun–Oct.
Habitat Open places in grasslands, pinelands
Elevation 10–1200 m [30–3900 ft]
Distribution
from USDA
North America; Mexico; South America (mostly Argentina)
[BONAP county map]
from FNA
AL; AR; FL; GA; KS; LA; MO; MS; NC; OK; SC; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Species 15–17 or more (11 in the flora).

Some horticultural gaillardias may be derived from hybrids (e.g., Gaillardia ×grandiflora van Houtte) involving G. aristata and some other species (probably G. pulchella). Such horticultural plants sometimes persist after cultivation or occur sporadically as waifs in places well beyond the “natural” ranges of the “parent” species; e.g., scattered localities in Arizona and California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plants with both ray and disc corollas wholly or mostly yellow to cream or white have been treated as a distinct species (Gaillardia lutea) or as varieties [G. aestivalis var. chrysantha (Small) Cronquist and G. aestivalis var. winkleri (Cory) B. L. Turner]. B. L. Turner (1979) formally recognized three varieties of G. aestivalis in the broad sense: var. aestivalis with purple discs, var. chrysantha with yellow discs, and var. winkleri with white discs. Although plants with yellow or white corollas may occur as local populations and may be distinguished by different ploidy levels, failing one or more additional distinguishing morphologic traits, they do not, in my opinion, merit formal taxonomic recognition.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves all basal (sometimes extending onto proximal 1/3+ of stems in desert annual G. arizonica), blades glabrous or sparsely villous (hairs jointed)
→ 2
1. Leaves basal and cauline or all cauline (restricted to proximal 1/3 or less of plants in some perennials; mostly cauline in annuals), blades hispidulous to scabrellous or strigillose to villous (hairs simple or jointed)
→ 3
2. Ray florets 0 or 7–10, corollas red to purple; disc corollas pinkish or purplish; pappus scales aristate, 6–9 mm
G. suavis
2. Ray florets 10–13, corollas yellow to orange; disc corollas yellow; pappus scales muticous or aristate, 2–5 mm
G. arizonica
3. Receptacles rarely setose (setae 0.1–1 mm); lobes of disc corollas attenuate-terete(bearing jointed hairs to 0.3 mm)
G. aestivalis
3. Receptacles usually setose (setae 0.3–6 mm); lobes of disc corollas ovate to deltate, sometimes attenuate (usually bearing jointed hairs 0.3+ mm, hairs shorter in G. coahuilensis)
→ 4
4. Annuals or perennials (seldom from woody caudices); cypselae hairy (hairs inserted at bases and sometimes on angles, not on faces)
→ 5
4. Perennials (often with woody caudices, seldom flowering first year); cypselae hairy (hairs inserted at bases and on angles and faces)
→ 8
5. Perennials (sometimes flowering first year); leaves basal and cauline; receptacular setae (2–)3–6+ mm
G. aristata
5. Annuals (seldom persisting); leaves all or mostly cauline; receptacular setae mostly 0.3–3 mm
→ 6
6. Hairs on disc corollas to 0.3 mm
G. coahuilensis
6. Hairs on disc corollas 0.3+ mm
→ 7
7. Ray corollas usually bicolored (brown-purple to red proximally, tipped with yellow or orange), rarely unicolored; lobes of disc corollas deltate to ovate, often attenuate, 1–3+ mm; cypselae all alike
G. pulchella
7. Ray corollas usually uniformly dark red to purple, rarely yellow; lobes of disc corollas mostly ovate-deltate to lanceolate, 0.5–1 mm; cypselae dimorphic (outer longer than inner and with shorter pappus scales)
G. amblyodon
8. Leaves (most or all) usually pinnatifid, rarely entire; disc corollas usually purplish or purple-tipped, rarely wholly yellow
G. pinnatifida
8. Leaves (most or all) entire or toothed; disc corollas yellow or purplish
→ 9
9. Leaf blades linear, 3–5(–8+) mm wide, faces sparsely hispidulous or glabrate; disc corollas usually purplish
G. multiceps
9. Leaf blades mostly oblanceolate to spatulate, usually (at least the larger) 10–25 mm wide, faces closely puberulent or sericeous to villous; disc corollas yellow
→ 10
10. Leaves mostly basal, sometimes some to mid stems; peduncles 10–30 cm; cypselae mostly 1–2 mm
G. parryi
10. Leaves mostly cauline; peduncles 2–6(–10) cm; cypselae mostly 3–4 mm
G. spathulata
Source FNA vol. 21, p. 421. Treatment author: John L. Strother. FNA vol. 21, p. 425. Treatment author: John L. Strother.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Gaillardiinae Asteraceae > tribe Heliantheae > subtribe Gaillardiinae > Gaillardia
Sibling taxa
G. amblyodon, G. aristata, G. arizonica, G. coahuilensis, G. multiceps, G. parryi, G. pinnatifida, G. pulchella, G. spathulata, G. suavis
Subordinate taxa
G. aestivalis, G. amblyodon, G. aristata, G. arizonica, G. coahuilensis, G. multiceps, G. parryi, G. pinnatifida, G. pulchella, G. spathulata, G. suavis
Synonyms Helenium aestivale, G. aestivalis var. flavovirens, G. chrysantha, G. fastigiata, G. lanceolata, G. lanceolata var. fastigiata, G. lanceolata var. flavovirens, G. lutea
Name authority Fougeroux: Observ. Phys. 29: 55. 1786 (as Gaillarda): Hist. Acad. Roy. Sci. Mém. Math. Phys. (Paris, 4to) 1786: 5. (1788) (Walter) H. Rock: Rhodora 58: 315. (1956)
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