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bonfire moss, common cord-moss, funaria moss

apiculate funaria moss

Habit Plants 4–10 or more mm, with a basal antheridial branch, medium green to yellowish green; leafless proximally with leaves crowded and bulbiform distally, sometimes laxly foliate throughout. Plants 3–5 mm, yellowish green, stem with leaves crowded at the tip forming a bulbiform spiral when dry and bearing a short antheridial branch at the base.
Leaves

smaller proximally, distal leaves 2–4 mm, deeply concave, oblong-ovate to broadly obovate distally, acute to apiculate or short-acuminate, entire or weakly serrulate distally;

costa subpercurrent to short-excurrent;

distal laminal cells thin-walled and inflated, hexagonal or oblong-hexagonal becoming much more oblong proximally.

few and reduced proximally, distal leaves 1.7–2.3 mm, concave, the blade ovate to obovate, abruptly contracted to a flexuose, hyaline tipped awn, distal blade margins weakly crenulate to rounded serrate above, entire proximally;

costa strong, excurrent with the distal end hyaline;

distal laminal cells thin-walled, rhomboid to elongate-hexagonal, proximal cells short-rectangular to almost quadrate toward the margins becoming oblong-rectangular near the costa.

Seta

usually (12–)20–45(–80) mm, slender and flexuose, usually hygroscopic.

usually 3–6 mm, flaring into a slender neck, hygroscopic.

Capsule

2–3.5 mm, pyriform, asymmetric, curved to straight, horizontal to pendent or merely inclined or nearly erect, becoming sulcate when dry below the strongly oblique mouth;

annulus revoluble, operculum slightly convex;

peristome brown, papillose-striate proximally and papillose distally, strongly trabeculate, becoming appendiculate distally, forming a lattice by fusion of the tips;

endostome segments lanceolate about 2/3 as long as the teeth, yellowish, finely papillose-striate.

1.5–2 mm, inclined to arcuate, asymmetric, obovoid, mouth oblique;

annulus none;

operculum conic-convex;

peristome inserted below the rim of the mouth;

exostome transparent and papillose, of low, truncate segments about 1/4 the length of the teeth, endostome teeth lanceolate, reddish with a hyaline tip, obliquely papillose-striate to the papillose hyaline tip, trabeculae weakly developed, appendiculae almost absent in the tip.

Calyptra

cucullate, smooth.

cucullate, smooth.

Spores

mostly 12–21 µm, finely papillose.

22–30 µm, nearly smooth to papillose.

Funaria hygrometrica

Funaria apiculatopilosa

Habitat Sand mixed with clay
Elevation moderate to high elevations
Distribution
map from USDA
Worldwide except Antarctica
[WildflowerSearch map]
map from FNA
TX; Mexico (Puebla, Zacatecas)
Discussion

Varieties ca. 20 (2 in the flora).

Funaria hygrometrica is one of the most common, weedy, and widely distributed mosses in the world; its distribution closely parallels that of Bryum argenteum. It is widely illustrated in textbooks to demonstrate the life cycle of a typical moss, possibly because of the abundant conspicuous sporophytes produced and its frequent presence in greenhouses. However, the peristome with opposite, instead of alternate, teeth in the two peristome rows is clearly atypical among the majority of mosses. Most of the varieties that have been described probably do not merit recognition because of the morphological plasticity of the species in response to environmental conditions.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Funaria apiculatopilosa is clearly distinct from other exannulate members of the genus in the very long-excurrent sinuose costa with a hyaline tip. The distal leaves spiral around the base of the seta when dry, and are spreading when moist. The trabeculae are thin, lightly pigmented and relatively few, and the spores are thin-walled. It is known from the Davis Mountains in Texas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Parent taxa Funariaceae > Funaria Funariaceae > Funaria
Sibling taxa
F. americana, F. apiculatopilosa, F. arctica, F. flavicans, F. microstoma, F. muhlenbergii, F. polaris, F. serrata
F. americana, F. arctica, F. flavicans, F. hygrometrica, F. microstoma, F. muhlenbergii, F. polaris, F. serrata
Subordinate taxa
F. hygrometrica var. calvescens, F. hygrometrica var. hygrometrica
Key
1. Capsule 2-3.5 mm, horizontal to pendent, curved, capsule neck less tapered, mouth of capsule narrow
var. hygrometrica
1. Capsule 2-3 mm, inclined to nearly erect, straight or weakly curved, capsule narrowly tapered to a long slender neck, mouth of capsule wide.
var. calvescens
Synonyms Entosthodon apiculatopilosus, F. orcuttii
Name authority Hedwig: Sp. Musc. Frond., 172. (1801) Cardot: Rev. Bryol. 40: 37. (1913)
Source FNA vol. 27, p. 190. Treatment authors: Donna H. Miller, Harvey A. Miller. FNA vol. 27, p. 193. Treatment authors: Donna H. Miller, Harvey A. Miller.
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