Froelichia gracilis
Amaranthaceae
slender cotton-weed, slender Snake-cotton
amaranth family
several (rarely 1), erect or ascending, sometimes procumbent, usually much-branched from base, slender, 1–5(–10) dm, densely or sparsely villous-tomentose with grayish white hairs.
without nodal spines (Amaranthus spinosus sometimes with paired nodal spines).
predominant on proximal 1/3 of plant, often basal, sessile;
blade linear to lanceolate or lance-elliptic, largest leaves 1.6–9(–13.5) × 0.2–0.9(–1.2) cm, base acuminate or attenuate, apex acute to acuminate, canescent or sericeous adaxially, sericeous-tomentose with white or gray hairs abaxially.
alternate or opposite, exstipulate, usually petiolate;
blade margins entire (entire or serrulate in Iresine; entire, crispate, or erose in Amaranthus).
cymules arranged in spikes, panicles, thyrses, heads, glomerules, clusters, or racemes; each flower subtended by 1 bract and 2 bracteoles (latter sometimes 1 or absent in Amaranthus).
sparsely branched, flowers arranged in 3-ranked spiral;
bracteoles stramineous or blackish, glabrous.
2.4–3.8 mm;
perianth lobes oblong-linear, apex acute or acutish, pubescence grayish;
filament lobes stramineous or blackish, blunt.
bisexual or unisexual (plants then monoecious or dioecious), hypogynous, generally small or minute;
tepals mostly (1–)4–5 or absent, distinct or connate into cups or tubes, scarious, chartaceous, membranaceous, or indurate;
stamens 2–5, filaments basally connate into cups or tubes, rarely distinct, alternating with pseudostaminodes (appendages on staminal tubes) or not, anthers 2-locular with 1 line of dehiscence or 4-locular with 2 lines of dehiscence;
ovary superior, 1-locular;
ovules 1 or, rarely, 2–many;
style 1 or absent;
stigmas 1–3(–5).
utricles, dry, dehiscent or not.
black, reddish brown, or brown, lenticular, subglobose or globose (rarely cylindric), usually small;
embryo peripheral, surrounding mealy perisperm.
2.5–5 × 2.7–4 mm, apex slightly oblique, with irregularly and deeply cut (“spiny”) lateral wings, both surfaces of perianth with distinct spines or tubercles.
= 54.
Froelichia gracilis
Amaranthaceae
The current range of Froelichia gracilis is due in large part to its introduction via railroads during the past 100 years, and the majority of records for the species east of the Mississippi River occur on or near railroads. S. F. Blake (1956) discussed this eastern spread of the species. Froelichia gracilis is considered a minor invasive weed in the Northeast; its adaptation to open sandy or gravely soils will restrict its spread to open sites with poor soil.
In regions where their ranges overlap, Froelichia gracilis may hybridize with F. floridana. Plants with intermediate form from Texas and the Midwest have been noted. Suspected hybrids appear closest in habit to F. gracilis and will generally key to that species. Floral structure will be intermediate and variable; the branching will be typically less than in F. gracilis and present distal to the base.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genera ca. 65, species ca. 900 (12 genera, 80 species in the flora).
Centers of diversity for Amaranthaceae are southwestern North America, Central America, South America, and Africa south of the Sahara Desert. Generic limits are not well defined in some groups; fewer than 60 or more than 70 genera could be recognized.
Some species occur in severe habitats such as sandy, calcareous, gypseous, saline, or serpentine soils in deserts, semideserts, and seashores. Some species are weedy, including the major agricultural weeds in Amaranthus. Some species are cultivated as ornamentals, particularly Amaranthus caudatus (love-lies-bleeding), A. hypochondriacus (prince’s-feather), A. tricolor (Joseph’s-coat), Celosia cristata (cockscomb), and Gomphrena globosa (globe-amaranth). Native Americans domesticated white-seeded grain amaranths (A. caudatus, A. cruentus, and A. hypochondriacus) for use as cereal grains. Some species of Amaranthus and Celosia are potherbs.
Amaranthaceae are usually divided into subfamilies Amaranthoideae (anthers 4-locular with two lines of dehiscence) and Gomphrenoideae Schinz (anthers 2-locular with one line of dehiscence). Amaranthaceae and Chenopodiaceae have long been recognized as allied families that share a number of features: generally small flowers, one perianth whorl, a syncarpous gynoecium with a superior ovary and often only one ovule, basal or free-central placentation, pollen characteristics, centrospermous embryo development, betalain pigments, and P-type form (c) sieve-element plastids.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Distal leaves alternate | → 2 |
1. Distal leaves opposite | → 5 |
2. Flowers unisexual (plants monoecious or dioecious); utricles 1-seeded | → 3 |
2. Flowers bisexual; utricles 2+-seeded | → 4 |
3. Shrubs | Iresine |
3. Herbs, annual, rarely perennial | Amaranthus |
4. Leaf blades mostly lanceolate, ovate, or deltate; pseudostaminodes absent | Celosia |
4. Leaf blades linear; pseudostaminodes alternating with filaments on staminal tubes | Hermbstaedtia |
5. Flowers unisexual (plants dioecious); inflorescences terminal, diffuse, open panicles | Iresine |
5. Flowers bisexual; inflorescences terminal and/or axillary glomerules, heads, or spikes | → 6 |
6. Inflorescences sessile glomerules or condensed spikes, axillary | → 7 |
6. Inflorescences pedunculate heads or spikes, terminal and sometimes axillary near stem tips | → 10 |
7. Inflorescences several-flowered, axillary spikes; pseudostaminodes present | Alternanthera |
7. Inflorescences 1-20-flowered, axillary glomerules; pseudostaminodes present or absent | → 8 |
8. Indumentum of branched or barbed trichomes, densely covering plant (rarely glabrous); pseudostaminoides present | Tidestromia |
8. Indumentum of simple trichomes; pseudostaminoides absent | → 9 |
9. Tepals connate proximally, tips 1-veined; filament tubes inserted distally on perianth tubes; basal rosette leaves usually absent at anthesis | Guilleminea |
9. Tepals distinct, 3-veined; filament tubes ± free from tepals; basal rosette leaves present at anthesis | Gossypianthus |
10. Inflorescences simple or compound spikes | → 11 |
10. Inflorescences globose or cylindric heads or spikes | → 12 |
11. Inflorescences mostly compound, interrupted spikes; tepals connate into indurate tubes with lateral crests or spines, lanate | Froelichia |
11. Inflorescences simple spikes or few-branched panicles, flowers progressively farther apart below; tepals basally connate into indurate tubes, without ornamentation, ± glabrous | Achyranthes |
12. Inflorescences not immediately subtended by leaves | Alternanthera |
12. Inflorescences immediately subtended by 2 or more leaves | → 13 |
13. Leaves fleshy, sessile; blade linear to narrowly obovate, glabrous except in axils | Blutaparon |
13. Leaves not fleshy, petiolate or sessile; blade ovate to obovate, pilose at least abaxially | Gomphrena |
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