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field snakecotton, Florida Snake-cotton, plains Snake-cotton

slender cotton-weed, slender Snake-cotton

Habit Plants annual; taproots semi-woody. Plants annual or short-lived perennial; taproots narrow, enlarged when perennial, semiwoody.
Stems

1, erect or ascending, sometimes procumbent, simple or sparsely branched (rarely much-branched) from base or above, stout, to 18 dm, puberulent or tomentulose with short, viscid, whitish or brownish hairs.

several (rarely 1), erect or ascending, sometimes procumbent, usually much-branched from base, slender, 1–5(–10) dm, densely or sparsely villous-tomentose with grayish white hairs.

Leaves

principally on proximal 1/3 of plant, petiolate;

blade lanceolate to oblanceolate, oblong, or linear, largest leaves 3.8–11.2(–21) × 0.5–3.8(–4.2) cm, base attenuate to cuneate, apex acute to obtuse, canescent to subscabrous adaxially, sericeous-tomentose abaxially.

predominant on proximal 1/3 of plant, often basal, sessile;

blade linear to lanceolate or lance-elliptic, largest leaves 1.6–9(–13.5) × 0.2–0.9(–1.2) cm, base acuminate or attenuate, apex acute to acuminate, canescent or sericeous adaxially, sericeous-tomentose with white or gray hairs abaxially.

Spikes

dense, much-branched, apex often pyramidal, flowers arranged in 5-ranked spiral;

bracteoles stramineous or blackish, pubescent with small tufts distally.

sparsely branched, flowers arranged in 3-ranked spiral;

bracteoles stramineous or blackish, glabrous.

Flowers

4–6 mm;

perianth lobes, greenish white to pinkish, oblong, apex acute;

filament lobes slightly to greatly recurved distally, stramineous to pinkish, apex acute.

2.4–3.8 mm;

perianth lobes oblong-linear, apex acute or acutish, pubescence grayish;

filament lobes stramineous or blackish, blunt.

Utricles

flask-shaped, 5 × 4–5 mm, with irregularly dentate lateral wings, both surfaces of perianth with distinct spines or tubercles.

2.5–5 × 2.7–4 mm, apex slightly oblique, with irregularly and deeply cut (“spiny”) lateral wings, both surfaces of perianth with distinct spines or tubercles.

2n

= 58 + 2.

= 54.

Froelichia floridana

Froelichia gracilis

Phenology Flowering summer. Flowering summer in north, year-round in extreme south.
Habitat Open sand prairies, edges of woodlands in sandy soils, roadsides, railroad rights-of-way Open plains, rocky hillsides, roadsides, waste ground, railroad ballast
Distribution
from FNA
AL; AR; CO; FL; GA; IA; IL; IN; KS; KY; LA; MD; MI; MN; MS; NC; ND; NE; OH; OK; SC; SD; TX; WI; West Indies [Introduced in Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; CA; CO; CT; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MO; MS; NC; NE; NH; NJ; NM; NY; OH; OK; PA; SC; TN; TX; VA; WI; WV; ON; Mexico (Chihuahua, Coahuila, Nuevo León, Tamaulipas); Asia (Japan) [Introduced in Europe]
[WildflowerSearch map]
[BONAP county map]
Discussion

Froelichia floridana ranges from broad-leaved stout herbs surpassing 1 meter in height to much-branched plants forming bushes to 2 meters in diameter and single-stemmed, erect herbs 1 dm in height. Much of the variation in the species has been difficult to ascribe to either geographic or ecologic differences and reflects inherent genetic variability and wide environmentally induced plasticity. Although specialists and taxonomists have implied the existence of well-delineated morphologic taxa with a strong geographic correspondence, this is not always true, and, in fact, one encounters typical specimens of any given variety sporadically throughout the range of the species.

Some variability in Froelichia floridana is clearly attributable to geography, principally in terms of general morphologic trends. There is a cline toward long, narrow, almost linear leaves as one moves east along the Gulf Coast and into Florida. This would culminate in the little-known var. pallescens Moquin-Tandon, a linear-leaved form (leaf length more than eight times width) from peninsular Florida. Additional variation ascribable to geography is observed in plants traditionally included in F. drummondii, these being large, stout plants with broader leaves, more obtuse leaf apices, and more densely fulvous pubescence on the abaxial surfaces of leaves. Plants of this form are generally restricted south of the Brazos River and continue to the southern extent of the range in Kenedy County, Texas. Further work examining this variation using micromorphology or molecular markers may elucidate a reliable means to identify intraspecific taxa within F. floridana.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The current range of Froelichia gracilis is due in large part to its introduction via railroads during the past 100 years, and the majority of records for the species east of the Mississippi River occur on or near railroads. S. F. Blake (1956) discussed this eastern spread of the species. Froelichia gracilis is considered a minor invasive weed in the Northeast; its adaptation to open sandy or gravely soils will restrict its spread to open sites with poor soil.

In regions where their ranges overlap, Froelichia gracilis may hybridize with F. floridana. Plants with intermediate form from Texas and the Midwest have been noted. Suspected hybrids appear closest in habit to F. gracilis and will generally key to that species. Floral structure will be intermediate and variable; the branching will be typically less than in F. gracilis and present distal to the base.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 4, p. 446. FNA vol. 4, p. 445.
Parent taxa Amaranthaceae > Froelichia Amaranthaceae > Froelichia
Sibling taxa
F. arizonica, F. drummondii, F. gracilis, F. texana
F. arizonica, F. drummondii, F. floridana, F. texana
Synonyms Oplotheca floridana, F. campestris, F. floridana var. campestris, F. floridana var. pallescens Oplotheca gracilis, F. braunii, F. texana, Oplotheca texana
Name authority (Nuttall) Moquin-Tandon: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 13(2): 420. (1849) (Hooker) Moquin-Tandon: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 13(2): 420. (1849)
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