Frasera speciosa |
Gentianaceae |
|||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
deer's-ears, elkweed, frasera, giant frasera, monument plant |
gentian family |
|||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habit | Herbs monocarpic, 5–20 dm, glabrous or stems and leaves puberulent. | Herbs [shrubs, trees], annual, biennial, or perennial, autotrophic, with green stems and leaves, or mycotrophic; when strongly mycotrophic, stems and leaves weakly chlorophyllous (only in Bartonia) or yellowish, whitish, purplish, or buff, or lacking chlorophyll (only in Voyria). | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | 1. |
|||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Leaves | blades not white-margined; basal spatulate or oblanceolate to elliptic-obovate, 7–50 × 1–15 cm, apex rounded to acute; cauline leaves whorled, blade oblong-lanceolate. |
cauline, often also basal, opposite, whorled, or rarely alternate, sessile or petiolate, simple; stipules absent [rarely present as ocreae]; blade margins entire. |
||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Inflorescences | elongate, open proximally, ± dense distally. |
cymes (sometimes racemoid, spicoid, or capitate), thyrses, or verticillasters, or solitary flowers; flowers pedicellate or sessile. |
||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Flowers | calyx 10–25 mm; corolla pale yellowish green, purple-dotted, occasionally suffused with purple distally, 12–25 mm, lobes elliptic-oblong to obovate, apex [obtuse or] acute to short-acuminate; androecial corona scales 7–9 mm, deeply multicleft; style slender, distinct; nectaries and foveae 2 per corolla lobe, foveae narrowly elliptic, opening directly adaxial to nectary, each opening with a ± even fringe all around, the pair opening into a green but not rimmed area on the corolla surface. |
bisexual or occasionally some unisexual [all unisexual on some or all plants], homostylous [heterostylous], protandrous and outbreeding or less often homogamous and autogamous, radially [somewhat bilaterally] symmetric, 4–12(–14)-merous [rarely 3-, 6-, or 16-merous] except for carpels; perianth hypogynous, calyx and usually corolla persistent; calyx green or occasionally ± hyaline (absent in Obolaria), sepals connate or some [or all] nearly distinct, lobes imbricate in bud, often ± unequal, colleters often present adaxially near base; corolla petaloid, petals connate, lobes contorted in bud or rarely imbricate (Obolaria, Voyria), spurs present only in Halenia, 1 per petal; stamens epipetalous, isomerous and alternate with petals, all fertile [rarely some sterile], equal [unequal]; filaments free or connected by a corona; anthers 2-locular, dehiscing longitudinally [with terminal pores], remaining straight, recurving, or coiling helically or circinately, distinct or (only in some spp. |
||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Fruits | capsular, dehiscence septicidal or rarely rupturing irregularly (Obolaria) [indehiscent capsules, berries]. |
|||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Seeds | few–very many, usually sessile; endosperm abundant and embryo small in autotrophic species, endosperm scant and embryo undifferentiated in completely mycotrophic species (Voyria). |
|||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Of | Gentiana) coherent; pistil 1, 2-carpellate; ovary 1[or 2]-locular; placentae 2, parietal [axile]; style present or absent, erect or initially deflexed to one side [declinate], uncleft, shallowly 2-cleft, or deeply cleft (Sabatia); stigmas 1 or 2, coiling only in Sabatia, decurrent on ovary (only in Lomatogonium, sometimes slightly so in Bartonia). |
|||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
2n | = 78. |
|||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Frasera speciosa |
Gentianaceae |
|||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Phenology | Flowering summer. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Open woods, montane to subalpine meadows. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 1500–3500 m. (4900–11500 ft.) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
AZ; CA; CO; ID; MT; NM; NV; OR; SD; TX; UT; WA; WY; Mexico (Nuevo León, Tamaulipas)
|
nearly worldwide |
||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Discussion | Swertia radiata var. maderensis Henrickson, endemic to Mexico, is conspecific with Frasera speciosa, but the necessary combination in Frasera has not been published. If that is done, our material will become var. speciosa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera ca. 100, species ca. 1800 (18 genera, 112 species in the flora). In the tribal classification by L. Struwe et al. (2002), genera 1–7 in this flora are in tribe Chironieae Dumortier, subtribe Chironiinae G. Don. Species of Chironieae generally lack nectaries, although Sabatia reportedly has indistinct nectaries at the base of the ovary. Genera 8–17 are in tribe Gentianeae Dumortier. Gentiana, in which the nectaries are on the gynophore, is in subtribe Gentianinae G. Don; the remaining genera of the Gentianeae in the flora area, all of which have epipetalous nectaries, are in subtribe Swertiinae Grisebach. Voyria constitutes the monogeneric tribe Voyrieae Gilg, in which the nectaries (when present) are on the ovary or the gynophore. Pedicel lengths given here refer to the true pedicels, between the most distal pair of bracts or bractlets and the calyx. In some genera, notably Centaurium, Sabatia, and Zeltnera, a flower terminating the ultimate branch of an inflorescence, directly subtended by bractlets, although sessile by this definition, may appear pedicellate. Corolla lengths as given are from the receptacle to the apices of the lobes (or plicae in Gentiana andrewsii). The Gentianaceae include many species esteemed in ornamental horticulture. In addition to those noted under the respective genera, the more important species in North American horticulture include Exacum affine Balfour f. ex Regel, Persian-violet, native to the island of Socotra, Yemen, which is widely grown as a florists’ pot plant. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Key |
|
|||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Source | FNA vol. 14. | FNA vol. 14. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Gentianaceae > Frasera | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Synonyms | F. macrophylla, Swertia radiata, Swertia radiata var. macrophylla, Tesseranthium radiatum, T. speciosum | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | Douglas ex Grisebach in W. J. Hooker: Fl. Bor.-Amer. 2: 66, plate 153. (1837) | Jussieu | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Web links |
|