Frangula alnus |
Frangula betulifolia |
|
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alder buckthorn, columnar buckthorn, European alder, European alder-buckthorn, fen buckthorn, glossy buckthorn, glossy false buckthorn, tall hedge buckthorn |
beechleaf frangula, birchleaf buckthorn |
|
Habit | Shrubs or small trees, 2–5(–7) m. Stems greenish to brown, sparsely puberulent, glabrescent. | Shrubs or small trees, 1–4 m. Stems brown to gray-brown, glabrous or pubescent. |
Leaves | deciduous; petiole 10–19 mm; blade dull green abaxially, glossy darker green adaxially, broadly elliptic-obovate to broadly elliptic or broadly oblong, 4–7(–11) cm, herbaceous, base cuneate to rounded, margins entire, apex rounded to acute or abruptly short-acuminate, abaxial surface sometimes minutely strigose along midveins, adaxial surface glabrous; secondary veins 6–10 pairs. |
deciduous; petiole (2–)5–16 mm; blade yellowish green abaxially, green adaxially, elliptic to oblong, elliptic-ovate or narrowly ovate, (4–)4.5–10 x (2–)2.5–5.5 cm, 1.6–2.6(–2.9) times longer than wide, ± herbaceous, base obtuse to truncate or rounded, margins serrate to subcrenate, apex usually acute to obtuse, sometimes slightly acuminate, both surfaces hirtellous, glabrescent; secondary veins (8–)9–13 pairs. |
Inflorescences | fascicles, sessile, 1–8-flowered. |
umbels, pedunculate, 2–20(–38)-flowered. |
Pedicels | 3–10 mm. |
3–7 mm. |
Drupes | black, globose or slightly elongate, 6–8 mm; stones 2(–3). |
black, globose, 5–10 mm; stones (2–)3(–4). |
Stigmas | 2–3-lobed. |
3-lobed. |
2n | = 20–26. |
|
Frangula alnus |
Frangula betulifolia |
|
Phenology | Flowering May–Sep. | Flowering Apr–Jun. |
Habitat | Alkaline and acid tamarack, red maple and cedar swamps, peatlands, bogs, fens, disturbed areas, riparian thickets, lakeshores, ditches, fencerows, hedgerows, low woods, beaver meadows. | Moist canyons, stream banks, rocky slopes, cliff bases, ledges, ridges, roadsides, deciduous, coniferous, and mixed woodlands. |
Elevation | 10–400 m. (0–1300 ft.) | 900–2800 m. (3000–9200 ft.) |
Distribution |
CO; CT; IA; ID; IL; IN; KY; MA; ME; MI; MN; NE; NH; NJ; NY; OH; PA; RI; TN; VT; WI; WV; WY; MB; NB; NS; ON; PE; QC; SK; Europe [Introduced in North America; introduced also in Asia (China, Russia), n Africa]
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AZ; NM; TX; Mexico (Chihuahua, Coahuila, Durango, Nuevo León, Sonora, Tamaulipas)
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Discussion | Two horticultural forms of Frangula alnus are widely sold and planted in North America. The cultivar 'Asplenifolia' (F. alnus var. asplenifolia Dippel; fern leaf or cutleaf buckthorn) has linear-oblong leaves with coarsely toothed to incised margins and commonly is grown as a specialty plant. The cultivar 'Columnaris' (tallhedge glossy buckthorn) has a narrow, upright habit and is used in hedging. Both forms were noted by M. H. Brand (http://www.hort.uconn.edu/plants/r/rhafra/rhafra1.html) to spread invasively by seeds. 'Asplenifolia' has been reported as naturalized in Ontario (A. W. Dugal 1989, 1992), Illinois (A. Branhagen, pers. comm.), and Ohio (M. K. Delong et al. 2005). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In the flora area, Frangula betulifolia is found in southeastern Arizona, the southern two-thirds of New Mexico, and trans-Pecos Texas. It and F. obovata are allopatric and morphologically distinct. C. B. Wolf (1938) considered a collection from Cochise County in southeastern Arizona, named as Rhamnus blumeri (Frangula ×blumeri), to be a hybrid between Frangula betulifolia and F. californica var. ursina; see the discussion of the latter taxon for more information. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 59. | FNA vol. 12, p. 58. |
Parent taxa | Rhamnaceae > Frangula | Rhamnaceae > Frangula |
Sibling taxa | ||
Synonyms | Rhamnus frangula | Rhamnus betulifolia |
Name authority | Miller: Gard. Dict. ed. 8, Frangula no. 1. (1768) | (Greene) Grubov: Trudy Bot. Inst. Akad. Nauk S.S.S.R., Ser. 1, Fl. Sist. Vyssh. Rast. 8: 268. (1949) |
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