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fissidens moss, yew-leaf pocket moss

Habit Plants to 11 × 3.5 mm. Plants to 7.5 × 1.5–2.5 mm.
Stem

mostly branched;

axillary hyaline nodules absent;

central strand present.

unbranched and branched;

axillary hyaline nodules absent;

central strand present or absent.

Leaves

as many as 15 pairs, often undulate, oblong to lanceolate, obtuse to broadly acute, cuspidate, to 2.7 mm × 0.6 mm;

dorsal lamina truncate-rounded proximally, ending at insertion, not decurrent;

vaginant laminae 2/3 the leaf length, ± equal, minor lamina ending on or near margin;

margin evenly serrulate, often crenulate-serrulate on dorsal and ventral laminae, elimbate;

costa ending in cuspidate apex, taxifolius-type;

laminal cells 1-stratose, distinct, smooth, bulging, firm-walled, irregularly hexagonal, 8–11 µm frequently paler at margin, conspicuously mammillose in vaginant laminae, larger and clearer juxtacostally.

as many as 25 pairs, often undulate, oblong-lingulate, abruptly narrowed to rounded-obtuse apex, 1.2–2 × 0.3–0.4 mm;

dorsal lamina narrowed proximally, rounded, ending a short distance before insertion;

vaginant laminae 1/2–2/3 the leaf length, acute, ± unequal, minor lamina ending near margin;

margin crenulate, elimbate;

costa ending 4–10 cells before apex, oblongifolius-type, distal part of leaf in transverse section showing 4 enlarged cells arranged in 2 rows;

laminal cells in dorsal and ventral laminae 1-stratose, distinct, lenticularly thickened but appearing mammillose, firm-walled, irregularly hexagonal, 6–9 µm, juxtacostal and interior cells of vaginant laminae 1-stratose, smooth, plane, larger;

medial marginal cells of vaginant laminae not differing perceptively from interior cells.

Seta

to 17 mm.

to 4.5 mm.

Sexual condition

rhizautoicous;

perigonia on short branches proximal to elongate stems;

perichaetia on short, proximal, axillary branches.

cladautoicous;

perigonia and perichaetia on main stems and branches.

Capsule

theca slightly inclined, slightly arcuate, bilaterally symmetric, to 1.5 mm;

peristome taxifolius-type;

operculum to 1 mm.

theca erect, radially symmetric, 0.3–1 mm;

peristome similiretis-type;

operculum 0.5–0.6 mm.

Calyptra

cucullate, smooth, to 2 mm.

0.7–0.8 mm.

Spores

13–18 µm.

11–15 µm.

Sporophytes

1 per perichaetium.

1 per perichaetium.

Fissidens taxifolius

Fissidens santa-clarensis

Habitat Damp, shaded soil, humus, and rocks Rotten wood, bark of living trees, particularly around the base, moist soil, and limestone
Distribution
from FNA
AL; AR; AZ; CA; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; ME; MI; MO; MS; NC; NE; NH; NJ; NY; OH; OK; OR; PA; SC; TN; TX; VA; VT; WA; WI; WV; NS; NT; ON; PE; QC; Mexico; Central America; West Indies; South America (Brazil and Chile); Europe; Asia (China, Japan); Pacific Islands (New Zealand)
[WildflowerSearch map]
from FNA
FL; Mexico; Central America; West Indies
Discussion

Fissidens taxifolius is distinguished by evenly serrulate or crenulate-serrulate leaf margin, stout costa ending in leaf cusp, conspicuous mammillose cells in the vaginant laminae, and sporophytes on short branches in the axils of proximal leaves. It can be confused with F. bushii, but that species is smaller, has costa shorter, and cells in the vaginant laminae have small, more or less inconspicuous papillae in the corners of the walls. Fissidens taxifolius usually occurs on shaded, damp soil or humus, whereas F. bushii is usually found on disturbed soil in woods, along paths, and in road cuts. Fissidens clebschii is considered to be a freakish expression of F. taxifolius (R. A. Pursell 2003).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Fissidens santa-clarensis, named for a province in Cuba, is restricted in the United States to peninsular Florida (Citrus, Collier, Dade, Levy, Manatee, Marion, Monroe, Polk, and Sumter counties). The species can be recognized by its abruptly narrowed, rounded-obtuse leaf apex, lenticularly-thickened dorsal and ventral laminal cells, short oblongifolius-type costa that in transverse section of the distal part shows four enlarged cells arranged in two rows, and similiretis-type of peristome. Collections of F. santa-clarensis were long assigned by American authors to F. radicans Montagne, a closely related species with caducous leaves, a broadly acute to rounded leaf apex, and an anomalous peristome with teeth undivided or irregularly divided.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 341. FNA vol. 27, p. 338.
Parent taxa Fissidentaceae > Fissidens Fissidentaceae > Fissidens
Sibling taxa
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. santa-clarensis, F. scalaris, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taylorii, F. ventricosus, F. zollingeri
F. adianthoides, F. amoenus, F. aphelotaxifolius, F. appalachensis, F. arcticus, F. asplenioides, F. bryoides, F. bushii, F. closteri, F. crispus, F. curvatus, F. dubius, F. elegans, F. exilis, F. fontanus, F. grandifrons, F. hallianus, F. hyalinus, F. leptophyllus, F. littlei, F. minutulus, F. obtusifolius, F. osmundioides, F. pallidinervis, F. pauperculus, F. pellucidus, F. polypodioides, F. scalaris, F. serratus, F. subbasilaris, F. sublimbatus, F. submarginatus, F. taxifolius, F. taylorii, F. ventricosus, F. zollingeri
Synonyms F. clebschii F. allenianus
Name authority Hedwig: Sp. Musc. Frond., 155, plate 39, figs. 1–5. (1801) Thériot: Mem. Soc. Cub. Hist. Nat. “Felipe Poey” 13: 209, plate 32, fig. 1. (1939)
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